lnoq€ puB Suol ulc (Z'g)q A-1.'I(90)'urals rad g g .r sa^eal auL[neJ.Buruel arll sE SuolsB lnoqB'po8ur,,\! alor]ad :Suuaqlr,r uoos 'Jalauerp urJ tI 01 dn go a11asoJ B SurutJoJ'pBorq urf, r'0 puB Suol urJ L 01 'a?Blnql€ds-alBFuola (02-0I) sa^aal IBrrp?U.so{rdsJo sasrqJpJ puE salcunpad saurocaq s^B,!\lB seJrdp asoq,$ guol rrJ q A ol slalqJuErq lua-puaDs€ 01. asonxau qlr^\ 'lJ€de urc g t-I slrxp JBal raddn uroq paqauBJq ,{l{?aal sualsra8uol:allasor oql Jo Japroq oql sprE^\ol urr gI-0I lnoq€ o1 ulc t ruorl qlSual urSurFuBJ (vI aJn8rd) suaurrJ?ds a3.re1 ur surals snoroJrrou :slrxB JEal pue sacrdB o1 pal-culsar ,{lalBurrlln urn?uaurnpur '5uol ulur 6 1 s.rreq pelurof poldurnJJ ol paAJnJ qlr,/r\paqlolc'a8B qlr.$ par qsru^\orq urnl qrII,$ sqrr qsrppar rBInFuE (,{llq8lls) q1r,tr uear8-qsrda.r8 '.{.uar '{ea.,!\ 'J€puals s({Ia?S olEr}sord Suruoraq uoos puB SurpBaJds uaql'lcaro

lsJU 1.8 '{crr{l ''urBrp ur IIIuI g 01 dn Jo looJ-dp} IEJlu") B urory Sursr.re (aJour Jo)0g 01 dn slooqs llqBr{ IeruuaJad E Jo aJuopr^a aArE lou op suaurrcads olqElre^€ aqJ

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:HJUgd :oloq) 'u s plalJuerJ 'f 'A'BL6I ^oN 6I ..'a1rq,r-{urd a{rds ra,\{old rur 0I 'qJaq (.{llPrpBr) SurpBards ̂lo.L, ErlBr}snv ura}sal!\ 'Japlnog 'peog eplequrey :snd,{";

'lrpaJaJ

racaoJpuE onblE unJolBdal urEncrdsuoc ur€JnlJnrls ,setruoc sBrlua)serolJur qo'lualuaquncord ulnlrqBq qo andrcaErd srldursap JnqpB snqaoads € urn^ou uoxBJ

'snlnJsnrsspJJ sr^aJq snl,{?s :urnJqEISatnrteto 'BrlunasueJl run8uol ureleu[uels urnqnl ur er^oJq eluaurvlLJ ',ErlIq.JaJeurur??s Jonlnt) 'ae)snI aeafcetq :sauBrsur (unroualxa urlresoBrd) runJosolrdurnJolBdal srlp^rnJar srlEluop suqBIS slprqle snqrcrdB ,rlnpur^ urap rarndJnd oruud'IlElnu€dur€Jqns 'rlJoJuoJ 'gi-gl saJoLr :urJ I x g I pB 'aBtruoJ(-r1e3uola) aErnlBuravrluaJsatolJuI €l€prdsnc'(BrluaJsaJr€ur €rorEur BlBlorlad-rlpl€ eSuol BrTeseq)urc a'0 x g'J p€ 'Blp^oqo alsn8u€ 16A plploaf,uPl(-u€aurl) BuJal[B eurlne) eL[ol'suassaJqBIS a^snurul snld anbruap Blnsrrq Jolr,\al ourud:srsourBJ a)rBd ,snqrluaqunc

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(A puE I sarn8rd) ..nou ds,lueg suaqunrord sntolrld .r

_ _ patEtsurar sr lueg (.32?rd A .r\ ) sn,{qreFfiuo{ .rE^ snfq)e1ltod d puE IuaB (iueB) Iluaalnd .rE^sn,{.Jrpls,tod d o}-luEr ur p€?npar sr IuaB rruaalnd a :,{aI e puE sarn'rl go pre aq1 q1r,$ parJrrElr puB passnt -srp s-r xalduo, sn,{qr?1s,qod .d ar{J .pepr^ord sr uaurcads odtr} eql go qde:r8oloqd e i.re,rioli aqj gb sfir.,reJpI€rrl,{lEuB ,{q palprlsnllr sr ,tcu?src lerru€loq arprESlooJ aqt !rr replnog rEau uro4 u^^oul ,{Fro.sarcads.{{all aqJ snloe?nl rE^ snrselor?ss A pue rorrul re^ sn?plnrrpradde d ,Frraqwnrortl a :possnrsrp puEpaqursap -3rB sn?olrld uerlDrlsny urelsaru Jo sartarre^ ,$au o$] pue sar:)ads .rau auo (!B6t) tlz tgz :€) t Ers?^nN Brlerlsny urels3^\ ur (aEarBqluerEuv) rg .u sntor?d uo salpnls rftuouoxEf c .lueg

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duEurrai) lsalU'6I uaqruntral 0008-(] '19 assErls JaturzueW 'ueqrunl\l SunltuIrIpssleetS aq)srueloB

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(r]ft6| tlz l:92:(g)t Drst^nN

264 Nuytsia Vol. 4, No. 3 (1983)

P : i l o a . s ! : o . ! d b . n s r . r l

r!1e!!

r o i l s l ] r . a d i r g l . r d i d : 1 1 ! : l r . r b .

. r o r . , s | r r h e t l n k - v h i r r .

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P r i l t t r t t r o c ! ( . . n s

Figure 1. Ptr'lotus procurabens Benl. A Holotype specimen (R. Cranfield s.n., PERTH). B-Part of ani.ot jpe sheer {M). (Photograph X. Liedl).

\at zt6Ez

pu" Suol tllur t I-A'I ol (C ,,{Z aJn8rd) aqnl snorqelS snoaceuBrqurau e 01 sasnurspaoJq r{?rd\ polrun sapourrus?s pu€ slua&relrd..Jaqlu€ drBluaurpnJ ,{J3A E qlrii\dllBuorsEcco Jo Suol rqul 9.0 o? xrpuaddp Ilpurs € q?ri uauo ,rauurql

JBrln^ouros ro 'adEqs puE ql8uol u! suaruBls slrual Jo sluauelrJ Eurllenba apourar"?s:uur 2.0 lnoqe01. po?plrp dll€seq 'a^oqe olelnqns .alppru

1B apr,r urur g0.0 .c ,Euol urur (1.0) 9:0ft'}) sluawellJ Jo lled aaJJ :lraJrad .{llualsrsuoJ suaurB?s rnod .r{lueuad uEql ra}roqsdlpa{Jeur wneooutS puu uruJaoJpuv .aqnl aql a^oq€ sur8ffur qloq .ro auo uo Bursr.repue Suol ruur I J osopou f11up3 padsr.rc srrBr{ aq1 ,elpprur

lnoqe la^ oo^\ flpuialur:rl?Bauaq pslJasur sJreq dq papaacxa lou lnq 3J€Jrns ralno uo poJncsqo leq,r -auros 'slBdal ralno ur upql ssal prer\lno ?u"q xacle alnJp .peorq urru 1,.0_9.0 (i O) pi e3uo1 urur (g't) I t-Z'g

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.Jsoprs qloq Jo ouo uo paaoJrBu dlldnlqe ssal Jo aroul puB alnlo^ur ,u,\\oJ8 dllnJ uaq,rapr,\! qIIu (T'I) 0'I-9 0 ot dn pue Suol urur (6.t) g.V-Z-t (CIZ arn3rd) sl"da? ra?no .xadEoql ldacxa asBJrns aql Sur,ra,roc urnlueurnpur ,8uol uIIu l.I o1 dn s.rreq Furpea.lds asop-ou-alellrcrlJa^qns lq8rBrls Sursr.rdruoc sledal Jo ocuaJseqnd lBsJoq

.1noq8no.rq1 elrqa,r-dro^r (urru 8'0-9 0 Jo) uorpod pcrde ,qsrlrq,ro Sururnl uaql 1sa5uo1 e3ur1 lurd Eurdajlanssrl snorrscs Surlnoqq8rau 'luocsaprrrl uaql rlurd o1 FurpBJ aldJnd pa8url EaJB uBrp-aur pPoJq :paJnoloc ,{luuoJrun lou sledal dn .raqSrq lua8.ra,tuoc ,BaJe snoocBuoo €EuuapJoq souo IBurFJBur

,aqnl aql a^oqe Iaa{ lure; Bub1 ruur Z-l.I B SutuuoJ urolprru 'sqrJ ploq sE aprslno podolo^ap sula^ aaJql:paAJncur ssal Jo aJour pu€ (Oa arnSrd)xad? aIrT-o8Bpuadde aql sprB,r\ol palrun ,{lpnsn sur8rBru snouBrqueur ,sa,qeq raOOnur i.lurpru aleqrul 'alpprtu o^oqB lsapeoJq ,Jrldrlla-.$orJEu o1 aleloacuel_.reaur1 s1eda"tr 'sl.JEJq .{q palBaruoJ lou oJaq.r sFdal Jo JnoloJ SurISElu lnq spJ€,!\dn ras.reds Bururor-aq arnlr?sa^ Iesrop :slalrrBq Suol Iuru t.I-9.0 Jo 8uu p .{q papunorrns .{lasuap ,Buolrurlr 8'0-1 0 lnoqp aqnl olEurqrnl B (sauo Jauur aql Jo ,,{llerJadsa) sluaurSas slr Jo sasEqpaua{Jrr{l. qlr^,r SuruJoJ 'suol ulur g .c Surqcea.r ,(sauo ralno eq1 3o fl.reurr.rd) qedelJo socrde ur.ru pasodxa Jo spJemlno Bur,r.rnc e q8norql ol€lnupduecqns ralei 1nqlJara ^llculs lsrrJ l€ 'pr8rr qjue1rod.a8e q1ral luacsauela ssal Jo e.rour ,dr1 Bursspd.rniJa^au'(CZ arnSrd) uorlrod IBJluao ur auJoq sJr€q ,!\aJ ,snoncrdsuoJur ura,tprur ,uor8a.r

IEcrdE alBururn)B pue u^rorq lq8q.ro; ldacxa snorlsnl .{ll€nsn ,1ue.redsuei1 pup utll 'apr^1 urrrl 8'I-t I '8uol ruru (g.g) S.g-g.Z (n.A) sapal)erg .lcodsB r{srdor5 o1 qsruaeJ8

IInp E a>lrds aloqM oql SuI^t8 dllpurJ rnoloc ?cBrq u,{orq ,(gZ arngrd) dll aq1 puo,{aq?€rlaauros ro ol Furlcaford sJrEq JJrls aql r{ll,r lnoqFnoJr{l ^lJPau aso|d dleleraporu 'uoruod

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Jo lJEd lpsBq puE slaJrpad Eurrnrsqo ,Euolurru A-8 0 puB lr{8rErls lsourlB sJr"q Je8uol ,dsr.rc sJrBq JalJoqs ,asopou-olplnJrluappue alrq,r sJrer{ aql'asoluaruol dlasuap ,Jalouerp urur 9.0-g.0,BBzErz ^llqErTS srqrpg. 'apt.u ruc 1 pue €uo1 urr g.I dllnJ 3ururu11e ,lerruoc-Buolqo q$ual 1B ,1yg e.rnSrilapvd urc 8'0-r'0'8uol ulc 0 I-9.0

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alrssasqns puE IEro1Bl'(I orn8rd) slolqcu€rq puB surals uo IBurruJal ,lJBduroJ ,^J€lrlos

s^€a,\lp'luBld {ad Og Ol 'J sa{ds .sruals roJ sE srr€q qlr^\ ol€1s gunod ur,{1uo sno[a. 'dJp uoq,r surS.reru uo alelnpun ualJo,saapal la.troy ur dlprcadsa pa8ur,r leq,r,radros

puB pouolleu 'ql8ua1 ur alqEuEA alor?ad :paJnoloc IJBp pup luaurctord aJEJrns ra,roluo qrJpplr'8uol urru 1

'c lurod luaunrxa u.rorq lq8g E qllr ,saceJJns qloq uo uaa.rfideop 'snooJeuoc .{llq8rTs 'a1B^oqo-,r\orJEu ol a}ploaJuBl( JEeurl) ,pBorq -- g-g.A

sn?olr?d luag c

266 Nuytsia Vol. 4, No. 3 (1983)

I'"

1r,"1 W''f,,,I

IruffiVH

,rrl

Figure 2. Ptrlot'us procumbens Benl. A-Spike. B-Biact, outer face. C Bracteole, outer face. D Outertepal, inner view. E-Inner tepal, inner vier,v. F-Androecium. C-Staminal tube opened. H Gynoecium.lDrawn from h,, l" trpe bv A. B-i thml.

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Iuag sn?auirpJ 'd' ' ' ' ' dol lB snollr^ ^JEAo :8urJ ea4 alnuru pqll.r unlcooJpuy 'sJrBq palelosr dq palsoJJ a8pr.r lensnun ue qll,t\lIEd Ja^lol ur slPdal 'ruJ

i x 0I ot dn sa^Bal aurlnB, 'lcara sluo?S Bu,uorq 'parrEq slJerg gv

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'lq8udn sulals aql pup paldnJJa?ur arB sa{rds aql orourJaqlJnJ'BrurJEI alnqrur B ol palroq€ sr uaurBls alrrals aql sarJads slr{} ur lnq,luag supue4al'd ur rncro osle suaruels 1ca;.rad ,rno; ^lluBlsuoC .ruc

a x gI o1 dn sa,real ,padeqs -JBa, qll& Ieruuarad lraro rre sr sarcods srql ,JaAa,roH .,,LvnrpLreuoqdrc wn!u!qc!JL,,,slarq

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sn?olr:td 'luag c

268 Nuytsia Vot. 4, No. 3 (1983)

Herb or subshrub with numerous branched prostrate stems forming close mats onground surface. More or less bushy plant with a persistent indumentum. Sfem c.18 cm long, branches and floriferous branchlets growing sympodially more or less atright angles (thus somewhat resembling P. obovatus (Gaudich) F. Muell.). Sp.ikeshemispherical or depressed-ovoid to 1.6 x 2.2 cm (hemispherical to elongate-ovoidand up to 4 x 2-6 cm in vat. appendiculatus). Bract 4.2-5.5 mm long, bracteoles 5-

l+ta:

t j l : rP , v , re - . r .

| ! ] . \ . t l ] . J , !un . ] t l l , r9

Figure 3. Ptilot'us appendiculatos Benl var. ninor Benl. Holotype specimen (S.-L. Everist 9195, BRI).(Photograph K. Liedl).

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2'tO Nultsia Vol. 4, No. 3 (1983)

times of the same colour. Outer tepals becoming 7.8 mm long and 2.5 mm wide; rnrertepals (woolly bearded inside) averaging 7 mm long and 1.8 mm broad, length of clawvery variable; stiff -harrs from initially purplish claw of bipartite tepals comparativelyshort, thus leaving major part of tepal Iimb uncovered, hairs never arising on limb.Filaments and. style to 3.2 mm long.

Other specimen examined. WESTERN AUSTRALIA: 6.4 km N of Mt Alice,Eremaean Province, "Basalt hill", N. I{. Speck 1185 (CANB, PERTH).

Distribution. Speck's two collections both came from the southern region of theAshburton Botanical District, Western Australia. These localities are about 400 kmfrom the Hamersley Range, the nearest area known for the type variety. Thus thereis no known overlap of ranges of the two varieties.

D-iscussion. At first glance the geographic separation and the significant differences(especially concerning colour of stems, leaves and bracts, size of floral organs as givenin Table 1) between the two infraspecific taxa seem to justify their recognition as twosubspecies. However, there is some variation in the colour of perianth and bracts andin the pubescence of tepals among the material of the type form (e.g. Parker 298, inAD, M, MEL, NT; Ashby 4166, in AD), which tend towards the corresponding fea-tures of var, luteoius. Therefore it seems more appropriate to give the new plant onlythe rank of varietas.

Table 1. Distinguishing morphological features between Ptilotus asttulasius var. ast'rolasius and var.

Character var. astrolasius var. luteolus

Colour of shootsColour of foliage

Indumentum ofyoung shoots and

Colour of spikes

Bracts andbracteoles

Outer tepalsInner tepalsClawLength of lilamentsand style

Creyish or yellowish turning to light brownHoary to greyish green turning to mid- andpale greenMealy tomentose with a close layer ofstellately branched hairs and fewer dendroid

Greenish to greyish-white with fadingreddish marksUsually inconspicuous, bracts almost colour-less, rarely brownish to deep brown, to 3 mmlong; bracteoles to 3.3 mm longTo 4.9 mm long, to 1.6 mm wideMean = 4.2 mm long, 1.1 mm wideSomewhat longer than limbTo 2.3 mm

YellowYellow

Dense villous pubescence of dendroidha1Is

Bicolorous with dark bract and yellowdistal parts of the tepalsMarkedly conspicuous by their dulldark colour; bract to 5 mm long;bracteoles to 6 mm longTo ?.8 mm long, to 2.5 mm wideMean = ? mm long, 1.8 mm wideVery va able in lengthTo 3.2 mm

4. Ptilotus polystachyus (Gaudich) F. Muell., Fragm. Phytogr. Austral.6:230(1868), emend. Benl, Mitt. Bot. MUnchen 3: 517 (1960)-Figure 4.

From the very beginning there has been some confusion regarding taxonomy andnomenclature of P. polystachyus. Gaudichaud's short diagnosis of Trichiniumpolystachyum from Shark Bay (in Freyc., Voy. Uranie: 445, 1826) did not clearlycharacterize the species. Mueller's description of Ptilotus polystachyus included T.sarlngii Lindley and ?. roseum Moq., both having been described in the meantime(see Bentham, FI. Austral. 5: 225, L870). Specimens of P. stirlingii at MEL were de-

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272 Nuytsia VoL 4, No. 3 (1983)

Figure 4. Pt'r?otus polyst'aclyus (Lindley) F. Muell. emend. Benl. A var. polystachyusi a-staminal cupspread open, inner view; b-ovary. B var. Iorgistac,\yus (W. Fitze.) Benl: a staminal cup spread open,innerview;b ovary. (Drawn from Hj. Eicller 18880 (A), Beauglehole 54011 (B)by A. Buhm).

NORTHERN TERRITORY: Simpson Desert (24'18'5, 136'37'E), P. K. Latz 4673(AD, CANB, NT).SOUTH AUSTRALIA: Western edge of Pernatty Lagoon (31'27'5, 137"11'E), H7.Eicfi.ler 18880 (AD, CANB, M).

4b. Ptilotus polystachyus var. polystachyus f. rubriflorus (J. M. Black) Benl,Mitt. Bot. Mijnchen 4: 282 (1962).NORTHERN TERRITORY: 40 miles (64 km) N of Wauchope Township, M.Lazarides 5843 (AD, CANB, M, NT, PERTH).SOUTH AUSTRALIA: Oodnadatta (27'33'3, 735"27'E), Nov. 1914, Mrss Staer s.n.(AD).

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214 Nuytsia Vot. 4, No. 3 (198A)

b. Stems and foliage remaining completely tomentose-pubescent. Re-corded from Western Australia between I7" and 22" lat . . . . . var. arthrotrichus(a) Fully developed spikes greenish turning brownish . . . . . 4c. f. arthrotrichus(b) Fully developed spikes red fading to brownish. . . . . - 4d,. f. ruber

3a. Spikes 1.6 2.3 cm across, with obviously narrowed apices unless fullygrown; bracts mostly acuminate, with (broad-) dark midrib,bracteoles to 5 x 3.5 mm. Perianth pink tipped, becoming creamy tobrown. . . . . . , , 4e. var , longis tachyus

b. Spikes 2.1 3 cm across, with roundish apices subconical when rmma-ture; bracts ovate-acute, usually with narrow rufous midribs,bracteoles to 4.5 x 3.5 mm. Perianth pale green becoming brownish

4f. var. pulleniiNore. Undoubtedly vat. arthrotichus is more closely related to var. polystachyusthan are var. longistachyus and var. pullenri. However, intermediates in floralcharacters between the latter taxa and the type variety, even within the samecollection (e.e. A. C. Beauglehole 5401?, R. Pullen 10669), forbid application of dif-ferent taxonomic ranks.

Acknowledgrnents

Mr A. B6hm, Mtinchen, prepared Figures 2 and 4, Mr K. Liedl, Munchen, pro-duced the photographs. Dr Hj. Eichler, Canbena, kindly read through taxa 2 and 3.Mr B. R. Maslln, Perth, and Dr A. Kanis, Canberra, gave useful suggestions toimprove the text. Dr J. W. Green, Western Australian Herbarium, and Dr R. W.Johnson, Queensland Herbarium, made available the specimens necessary to thestudies. The author wishes to express his deepest thanks to all these gentlemen.