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Vegetative Morphology

New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

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Page 1: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Vegetative Morphology

Page 2: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

How would you describe these plants?

Page 3: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM
Page 4: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM
Page 5: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Morphological description에 대한 용어는

1) 교과서1) 교과서

2) 웹페이지의 “Photo Gallary of Vascular Plant and Terminology”

h // b ll /Ph G ll OfV l Pl /h l/h l h lhttp://amborella.net/PhotoGallaryOfVascularPlants/html/help.html

3) “이창복도감 용어도해” 를 참고할 것

Page 6: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Pl t H bitPlant Habit 습성

· HERB 초본 – no above ground persistent woody tissue but may have underground perennating structures; may be annual 일년생, g p g ybiennial 이년생, or perennial 다년생

· SUBSHRUB 아관목 (=suffrutescent) – lower stems woody but upper stems herbaceous (prefix "sub" means "almost")stems herbaceous (prefix sub means almost )

· SHRUB 관목 – a woody low-stature perennial plant with one to many slender trunks arising from near its base

· TREE 교목 – a large woody perennial plant with one to several relatively massive trunks and an elevated crown

· SUCCULENT 다육성 – possessing thick usually soft watery leaves· SUCCULENT 다육성 – possessing thick, usually soft, watery leaves and/or stems. Stem succulents & leaf succulents

· VINE 초본성 덩굴식물 – a woody or herbaceous plant with a long, f fslender, more or less flexible stem which cannot support itself

· LIANA 목본성 덩굴식물(만목) – a woody, climbing vine (characteristic of the tropics)tropics)

Page 7: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM
Page 8: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

LianaLiana

vein

Page 9: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Roots 뿌리

• usually the portion of the plant that absorbs water and minerals

• mostly underground• mostly underground • principal organ of attachment • lacks nodes and buds (a node is that portion ( p

of the stem where the leaves originate) Stems 줄기

l f b i i i f l• leaf-bearing main axis of a plant • aerial or subterranean • divided into nodes and internodesdivided into nodes and internodes • gives rise to branches, leaves, and flowers Leaves 잎• lateral appendages on a stem • usually serves as the primary photosynthetic

surface of the plantsurface of the plant • can be extremely modified in morphology

Page 10: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Root typesTAPROOT t l i· TAPROOT 주근 – central main root that descends vertically; larger than any branching root

· FIBROUS 수염뿌리 – thin thread-like roots arising from a taproot or from stem tissuetaproot or from stem tissue

· ADVENTITIOUS 부정근 – roots that originate from any part of th l t th th th tthe plant other than the root system

Question: 왜 rhizome은 지하”경” 즉 줄기 이고 수평으로 뻗어나가는뿌리라고 할 수 없는가?

Page 11: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Stem types

· RHIZOME 지하경 – an underground horizontal persistent stem; its leaves are often reduced to scales and it usually

구경

ybears adventitious roots and buds

· STOLON 포복경 – a horizontal stem near the d f h d lground surface that produces new plants

at its nodes or tip (runner)

· BULB 인경 – a thickened, underground BULB 인경 a thickened, underground short vertical stem with large storage leaves; usually borne below ground

지하경괴경

· CORM 구경 – a solid, erect underground stem with leaves absent or dry and scale-like

포복경like

· TUBER 괴경 – a solid, enlarged, underground horizontal stem that serves

지하경

as a storage area for food reserves 포복경

Page 12: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

포복경

StolonStolon (=Runner)(=Runner)

포복경

괴경

bulb

corm인경

구경 지하경

Page 13: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Stem features

· NODE 절 – the position on a stem where a leaf or NODE 절 the position on a stem where a leaf or bud is or was attached

· INTERNODE 절간 – the portion of a stem between two nodes 정아

· AXIL 잎겨드랑이; 액 – the upper angle between a leaf (or any other lateral structure) and the stem to which it is attached

아린

· AXILLARY BUD 액아 – a bud borne in the axil of a leaf (also called a lateral bud)

· BUD 눈; 아 – the structure giving rise to a leafy

측아

피목엽흔

액아

stem, a flower, or both; it may be naked or protected by bud scales or stipules; it may be lateral or terminal

BUD SCALES 아린 l lik l th t t t

절간

· BUD SCALES 아린 – scale-like leaves that protect the buds

· BUD SCALE SCARS 아린흔 – scars remaining when the bud scales fall off

관속흑

the bud scales fall off

· STIPULES 탁엽 – usually a pair of appendages located at the base of a leaf but may be fused into a ring around the stem; variable in size,

g ; ,shape and texture; serves for protection or to attract pollinators

탁엽

Page 14: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

· STIPULE SCARS 탁엽흔 – a pair of scars or pa single ring-like scar when stipules fall away

· LEAF SCAR 엽흔 – the scar left when a leaf falls from a twig; it contains oneleaf falls from a twig; it contains one or more vascular bundle scars

Page 15: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

잎 (leaf) = 엽신 + 엽병 + 탁엽

엽맥

윗면; 배면; 등; 향축면줄기

싹; 芽(측아) 윗면; 배면; 등; 향축면아랫면; 복면; 배

엽연엽병엽병

탁엽

엽선엽신cf. pulvinus: 비대부위

Page 16: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Leaf persistence · EVERGREEN 상록성 – bearing green foliage all year round· EVERGREEN 상록성 – bearing green foliage all year round · DECIDUOUS 낙엽성 – the loss of leaves during unfavorable

conditions (such as at the end of each growing season)

Leaf parts · LEAF BLADE 엽신 – expanded portion of a leaf 엽신 p p· PETIOLE 엽병 – stalk of a leaf · STIPULES 탁엽 – the pair of appendages located at the base

of a petioleof a petiole

Page 17: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM
Page 18: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Leaf attachment

· PETIOLATE 엽병을 갖고 있는 – having a petiole

· SESSILE 엽병이 없는 – lacking a petiole (or leaf stalk)

· SHEATHING 엽초를 갖는 – leaf base enwraps stem

Leaf arrangement 엽서Leaf arrangement 엽서

- ALTERNATE 호생 – one leaf per node

- OPPOSITE 대생 – two leaves per nodeOPPOSITE 대생 two leaves per node

- WHORLED 윤생 – three or more leaves per node

Page 19: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

호생 대생 윤생

※ 참조: 90도씩 돌려가며 대생하는것※ 참조: 90도씩 돌려가며 대생하는것교호대생(decussate)

Page 20: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Leaf complexity

· SIMPLE LEAF 단엽 – a leaf with a single blade (it is not di id d i l fl h i l fl f bl ddivided into leaflets; there is always a flange of blade tissue connecting adjacent lobes)

· COMPOUND LEAF 복엽 – a leaf with more than one blade l d f l fl dper petiole (it is made up of two or more leaflets and

these leaflets are wholly separate)

- PETIOLULE 소엽병 – the stalk of a leaflet

- LEAFLET 소엽 – one of the segments of a compound leaf

- RACHIS – the axis of a pinnately compound leaf

· PINNATELY COMPOUND 우상복엽 – a leaf in which there 우상복엽

are more than three leaflets arranged in two rows along a common axis. The leaflets are attached like the vanes of a feather

· PINNA (pl. PINNAE) – the primary division (or leaflet) of a pinnately compound leaf

· BIPINNATELY COMPOUND 2회 우상복엽 – leaf divided twice BIPINNATELY COMPOUND 2회 우상복엽 leaf divided twice

· TRIPINNATELY COMPOUND 3회우상복엽 – leaf divided three times; a pinnule is the ultimate division of a 2- or 3-times compound leafcompound leaf

Page 21: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Tripinnately compound leaf

Bipinnately compound leaf

Page 22: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

· PALMATELY COMPOUND 장상복엽 –where the leaflets arise from a

i f h hcommon point of attachment; there is no rachis

· TRIFOLIOLATE 3출엽 – a compound leaf with three leafletsleaf with three leaflets

※ TRIFOLIATE 잎이 3개나는 윤생– three separate leaves arising from the same nodesame node

Simple leaf

Simple leaf or compound leaf?

Compound leafCompound leaf

Page 23: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

우수우상복엽 기수우상복엽

2회기수우상복엽

장상복엽 삼출엽 단신복엽

Page 24: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Leaf venationLeaf venation · PINNATE 우상맥 – consisting of a central mid-

vein with many secondary veins emerging on b th id t f f th lik ttboth sides to form a feather-like pattern

· PALMATE 장상맥 – all primary veins arise at the same point at the base of the leaf

· PARALLEL 평행맥 – veins lie more or less parallel to the leaf margins

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Leaf blade shape · LINEAR 선형 – long and narrow with the sides

parallel (>4:1)parallel (>4:1) · OBLONG 장타원형 – nearly rectangular with the

sides parallel (2-4:1) · LANCEOLATE 창형 – spear-shaped; widening LANCEOLATE 창형 spear shaped; widening

above base and then long tapering to apex (3-4:1)

· OVATE 난형 – egg-shaped; broad nearest base OVATE 난형 egg shaped; broad nearest base (<3:1)

· OBOVATE 도란형 – ovate, but with narrower end towards point of attachmenttowards point of attachment HINT: The prefix "ob" means opposite, so "obovate" is the opposite of "ovate."

· ELLIPTIC 타원형 – ellipse-shaped; widest near ELLIPTIC 타원형 ellipse shaped; widest near middle and tapering at both ends

· ORBICULAR 원형 – circle shaped · RENIFORM 신장형 – kidney shaped RENIFORM 신장형 kidney shaped

Page 28: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM
Page 29: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Heterophylly (이형엽성): 동일한 식물체에서도 다른 모양의 잎을 갖는 경우

Page 30: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Leaf apices 엽두

· ACUTE 예두 – sharp ending in a point with ACUTE 예두 sharp, ending in a point with straight sides to the apex (<90 degrees)

· ACUMINATE 점첨두 – sharp, ending in a long-tapering point with concave sides

· OBTUSE 둔두 – blunt, rounded (>90 degrees)

MUCRONATE 급첨두 ll b t i t· MUCRONATE 급첨두 – a small, abrupt point

Leaf bases 엽저· ACUTE 예저 – sharp <90 degrees ACUTE 예저 sharp, <90 degrees · ACUMINATE 유저 – sharp, long-

tapering point · OBTUSE 둔저 – blunt >90 degrees OBTUSE 둔저 blunt, >90 degrees · CORDATE 심장저 – heart shaped (equal

rounded lobes at the base) · OBLIQUE 왜저 – unequal sized lobes at OBLIQUE 왜저 unequal sized lobes at

base· PELTATE 순저 – umbrella like; the petiole

is attached to the blade insideis attached to the blade inside of the margin; often orbicular in shape

Page 31: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

예두 둔두 둔두

요두 절두

유두원두급첨두

Page 32: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

예저 둔저 설저

원저 유저 절저원저 유저 절저

심장저 열저

화살저

Page 33: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Leaf margins 엽연 거치· ENTIRE 전연– a margin without any toothing or division (smooth) · CRENATE 둔거치 scalloped or round toothed· CRENATE 둔거치 – scalloped or round-toothed · SERRATE 예거치 – a saw-toothed margin with sharp teeth pointing towards

the apex DENTATE 치아상거치 h h j i i h l f h i· DENTATE 치아상거치 – sharp teeth projecting at right angles from the margin

· PINNATELY LOBED 우상열 – lobed towards the midrib but not reaching it · PALMATELY LOBED 장상열 – lobes all arising from one point at the base of g p

the leaf

Page 34: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM
Page 35: New Vegetative Morphology - Amborella · 2011. 4. 8. · Microsoft PowerPoint - Week06-Morphology-vegetative Author: Administrator Created Date: 4/8/2011 9:23:20 PM

Surface features · GLABROUS 털이 없는 – lacking hairs

(trichomes); a smooth surface(trichomes); a smooth surface · PUBESCENT 털이 있는 – covered with

hairs (also called trichomes) STELLATE 성 가 있는 h i h· STELLATE 성모가 있는 – hairs that branch at or near their base (star-shaped from above)

· GLANDULAR 선모가 있는 – hairs that bear glands (that break down into sticky beads of fluid); they may be t lk d ( ti it t ) ilstalked (stipitate) or sessile

※ trichome: 털, 모용 = indumentum

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Special features· TENDRIL 덩굴손 – an

elongated twining segmentelongated, twining segment of a leaf, stem, or inflorescence by which a plant clings to its supportclings to its support

· THORN 경침 – a woody, sharp-pointed, modified stem (has stem like vasculature)stem-like vasculature)

· SPINE 엽침 – a sharp-pointed modified leaf or leaf part STIPULAR SPINE 탁엽성 엽침 borne- STIPULAR SPINE 탁엽성 엽침 – borne in pairs and lateral to leaf (or leaf scar)

· PRICKLE 피침 – a sharp pointed outgrowth from thepointed outgrowth from the epidermis