Plant Megafossils 9

Plant Megafossils

Kazuhiko Uemura1, Harufumi Nishida2, Mitsuo Suzuki3, and Arata Momohara4

1Department of Geology and Paleontology, National

Science Museum, Tokyo, 169-00073 Japan 2Faculty of Science and Engineering, Chuo University,

Tokyo, 112-8551 Japan 3Botanical Gardens, Graduate School of Science, Tohoku

University, Sendai, 980-0862 Japan 4Faculty of Horticulture, Chiba University,

Matsudo, 271-8510 Japan * indicates that the species has a nomenclatural problem, mostly in the typification (Art. 37) of ICBN Abies aburaensis Tanai, 1961 Jour. Fac. Sci., Hokkaido Univ., Ser. 4, vol. 11, p. 247, pl. 1, fig. 8 Holotype: UHR-15064 [NSM-PP] Abura, Setana-cho, Setana-gun, Hokkaido Kunnui Formation Middle Miocene [late Early Miocene] Abies firmoides Watari, 1956 Jour. Fac. Sci., Univ. Tokyo, Sec. 3, vol. 6, p. 427, photo. 4; text-fig. 4 Holotype: Bot. Inst., Fac. Sci., Univ. Tokyo, no. 31602 [TI] Kuji district, Iwate Prefecture [Noda Group] Early Oligocene Abies hokiensis Ozaki, 1979 Sci. Rep. Yokohama Natn. Univ. Sec. 2, no. 26, p. 35, pl. 1, fig. 8 Holotype: NSM-PP 16219 Tatsumi-toge, Saji-son, Yazu-gun, Tottori Prefecture Tochiwara Formation Late Miocene Abies honshuensis Tanai, 1961 Jour. Fac. Sci., Hokkaido Univ., Ser. 4, vol. 11, p. 247, pl. 1, fig. 2 Holotype: GSJ-4210 Aburato coal mine, Kamo-machi, Tsuruoka City, Yamagata Prefecture Aburato (coal-bearing) Formation Early Miocene For many helps and comments in the present work, we thank following persons: T. Asakawa, M. Masuda, T. Ohana, K. Saiki, Y. Shicchi, K. Terada, M. Tsukagoshi, and A. Yabe

Abies n-suzukii Tanai, 1961 Jour. Fac. Sci., Hokkaido Univ., Ser. 4, vol. 11, p. 249, pl. 1, fig. 13 Holotype: UHR-15067 [NSM-PP] Yoshioka, Fukushima-cho, Matsumae-gun, Hokkaido Yoshioka Formation Middle Miocene [late Early Miocene] Abies protofirma Tanai, 1961 Jour. Fac. Sci., Hokkaido Univ., Ser. 4, vol. 11, p. 248, pl. 1, fig. 10 Holotype: GSJ-4059 Onbara, Kamisaibara-son, Tomata-gun, Okayama Prefecture Onbara Formation Mio-Pliocene Abies sanzugawaensis Huzioka et Uemura, 1974 Bull. Natn. Sci. Mus., vol. 17, p. 343, pl. 1, figs. 1, 1a, b Holotype: AKMG-7470 Shimoshinden, Takamatsu, Yuzawa City, Akita Prefecture Sanzugawa Formation Late Miocene Abies ugoensis Huzioka et Uemura, 1973 Bull. Natn. Sci. Mus., vol. 16, p. 697, pl. 1, figs. 1, 1a Holotype: AKMG-7277 Kamitozawa, Nishiki-mura, Senboku-gun, Akita Prefecture Miyata Formation Late Miocene Abiocaulis yezoensis Suzuki, 1910 Bot. Mag. Tokyo, vol. 24, no. 284, p. 181, pl. 7, photos 1-2 Holotype: Bot. Inst., Univ. Tokyo [TI] Ikushunbetsu, Hokkaido Late Cretaceous Acer akagawaensis K. Suzuki, 1959 Monogr. Assoc. Geol. Collab. Japan, no. 9, p. 41, pl. 5, fig. 2 Holotype: IGF-1162 [FM] Loc. Ak1, Tennoji, Iizaka-machi, Shinobu-gun, Fukushima Prefecture Tennoji Formation Late Miocene Acer chiharae Huzioka et Nishida in Huzioka, 1964 Jour. Min. Coll. Akita Univ., Ser. A, vol. 3, no. 4, p. 88, pl. 14, fig. 5 Holotype: AKMG-3312 Seki, Aikawa-machi, Sado-gun, Niigata Prefecture Suginoura Formation Early Miocene (The species was described by Huzioka and Nishida (1960, Publ. Sado, Mus., no. 3, p. 18, pl. 5, figs. 6-8) without typification, and validly published in Huzioka (1964))

10 K. Uemura, H. Nishida, M. Suzuki and A. Momohara

Acer cuneatum Huzioka, 1972 Jour. Min. Coll. Akita Univ., Ser. A, vol. 5, p. 62, pl. 8, fig. 1 Holotype: IGPS-92366 Yongpukdong, Hamgyeong-bukdo, Korea Engelhardtia bed Middle Miocene [Oligocene] Acer debilum Huzioka et K. Suzuki, 1954 Trans. Proc. Palaeont. Soc. Japan, N.S., no. 14, p.140, pl. 16, figs. 11-12 Syntypes: Inst. Earth Sci., Dept. Arts Sci., Fukushima Univ. [FM] 1 km west of Mt. Tate-yama, Kitakata City, Fukushima Prefecture Shiotsubo Formation Late Miocene Acer endoanum Huzioka, 1972 Jour. Min. Coll. Akita Univ., Ser. A, vol. 5, p. 63, pl. 14, fig. 5H Holotype: Dept. Geol. Mineral., Hokkaido Univ. (Huzioka, 1943, pl. 22, fig. 6; Acer sp.) Hamjindong, Hamgyeong-bukdo, Korea Hamjindong Formation, Myeoncheon Group Middle Miocene Acer ezoanum Oishi et Huzioka, 1943 Jour. Fac. Sci., Hokkaido Imp Univ., Ser. 4, vol. 7, no. 1, p. 89, pl. 11, figs. 1-4 Syntypes: Dept. Geol. Mineral., Hokkaido Univ., nos. 9359a-b, 9360 Shiretori coal-mine, Shiretori-machi, Shiretori-gun, Saghalien [Makarov, Sakhalin, Russia] Naihoro (coal-bearing) Formation Middle Miocene [late Early Miocene] (Lectotype: no. 9360, by Tanai, 1983 in Jour. Fac. Sci., Hokkaido Univ., Ser. 4, vol. 20, p. 320) Acer fatsiaefolia Huzioka, 1943 Jour. Fac. Sci., Hokkaido Imp. Univ., Ser. 4, vol. 7, no. 1, p.131, pl. 23, fig. 7 Holotype: Dept. Geol. Mineral., Hokkaido Univ. Kinkodo, Usen-men, Geizitsu-gun, N. Keisyodo, Korea Ennichi Series Miocene Acer fujitogensis K. Suzuki, 1961 Sci. Rep. Fukushima Univ., no. 10, p. 83, pl. 18, fig. 5 Holotype: IGF-5191 [FM] Az-28c, Fuji-toge, Yanaizu-machi, Kawanuma-gun, Fukushima Prefecture (373344N, 1394322E) Lower part of Fujitoge Formation Late Miocene

Acer huziokae Tanai, 1983 Jour. Fac. Sci., Hokkaido Univ., Ser. 4, vol. 20, no. 4, p. 326, pl. 11, fig. 2 Holotype: TPM-400 (Tottori Pref. Mus.) Tatsumitoge, Saji-son, Yazu-gun, Tottori Prefecture Tatsumitoge Formation Late Miocene Acer imaii Huzioka et Nishida in Huzioka, 1964 Jour. Min. Coll. Akita Univ., Ser. A, vol. 3, no. 4, p. 90, pl. 14, fig. 11 Holotype: AKMG-3317 Seki, Aikawa-machi, Sado-gun, Niigata Prefecture Suginoura Formation Early Miocene (The species was described by Huzioka and Nishida (1960, Publ. Sado, Mus., no. 3, p. 18, pl. 6, figs. 1, 2) without typification, and validly published in Huzioka (1964)) Acer ishikariense Tanai, 1983 Jour. Fac. Sci., Hokkaido Univ., Ser. 4, vol. 20, no. 4, p. 322, pl. 5, fig. 8 Holotype: HUMP-26209 [NSM-PP] Reisui-zan, Yubari City, Hokkaido Ikushunbetsu Formation Early Oligocene [late Middle Eocene] Acer iwaii K. Suzuki, 1961 Sci. Rep. Fukushima Univ., no. 10, p. 81, pl. 18, figs. 1-4 Syntypes: IGPS-78144, 78141, 78143, 78142 IGPS loc. no. Fs-62 (loc. Jo-11), north cliff of northwestern valley of Shichiku, Yotsukura-machi, Futaba-gun, Fukushima Prefecture (37810N, 1405415E) Shichiku Formation Early Miocene Acer iwatense (Watari) Watari, 1952 see Acerinium iwatense Watari, 1941 Acer kokangenense Endo, 1950 Short Pap. IGPS, no. 1, p. 14, pl. 3, fig. 10 Holotype: IGPS-60993 Ryuhoku-do, Kokangen, Keigen-gun, Kankyo-hokudo, North Korea Engelhardtia bed Miocene [Oligocene] Acer konnoi Murai, 1963 Tech. Rep. Iwate Univ., vol. 16, no. 1, p. 108, pl. 13, fig. 4 Holotype: IAGI-61096 Yoake-zawa Ma-5, east of Oshuku hot-spring, Shizukuishi-cho, Iwate-gun, Iwate Prefecture (3937568N, 14056254E) Masuzawa Formation

Plant Megafossils 11

Late Miocene Acer koreanicum Endo, 1950 Short Pap. IGPS, no. 1, p. 15, pl. 3, fig. 13 Holotype: IGPS-60996 Ryuhoku-do, Kokangen, Keigen-gun, Kankyo-hokudo, North Korea Engelhardtia bed Miocene [Oligocene] Acer kushiroanum Tanai, 1970 Jour. Fac. Sci., Hokkaido Univ. Ser. 4, vol. 14, p. 489, pl. 20, fig. 1 Holotype: HUMP-25990 [NSM-PP] Nakanosawa, near Yubetsu coal mine, Yubetsu, Akan-machi, Akan-gun, Hokkaido Yubetsu Formation Early Oligocene [Late Eocene] Acer macrosamarum Huzioka, 1964 Jour. Min. Coll. Akita Univ., Ser. A, vol. 3, no. 4, p. 90, pl. 14, fig. 12 Holotype: AKMG-3933 Arase coal-mine, Tsuyukuma, Ani-machi, Kita-Akita-gun, Akita Prefecture Aniai (coal-bearing) Formation Early Miocene Acer matsuii Tanai et Onoe, 1959 Bull. Geol. Surv. Japan, vol. 10, no. 4, p. 282, pl. 6, fig. 1 Holotype: GSJ-4042 Shichiku, Ono-mura, Iwaki-gun, Fukushima Prefecture Basal part of Yunagaya Group [Shichiku Formation] Early Miocene Acer megasamarum Tanai et N. Suzuki, 1960 Jour. Fac. Sci., Hokkaido Univ., Ser. 4, vol. 10, p. 560, pl. 5, fig. 3 Holotype: UHR-15027 [NSM-PP] Abura, Setana-cho, Setana-gun, Hokkaido Kunnui Formation Middle Miocene [late Early Miocene] Acer meisenense Endo, 1950 Short Pap. IGPS, no. 1, p. 12, pl. 3, fig. 4 Holotype: IGPS-60990 Kanchindo, Meisen-gun, Kankyo-hokudo, North Korea Kanchindo Formation Miocene [Middle Miocene] Acer miojaponicum Huzioka et Nishida, 1960 Publ. Sado Mus., no. 3, p. 19, pl. 5, fig. 5. Holotype: [AKMG] Seki, Aikawa-machi, Sado-gun, Niigata Prefecture

Suginoura Formation Early Miocene (See also Huzioka (1964, Jour. Min. Coll. Akita Univ., ser. A, vol. 3, no. 4, p. 92) under A. protojaponicum Tanai et Onoe) Acer miyagiense Endo, 1950 Short Pap. IGPS, no. 1, p. 16, pl. 3, fig. 14 Holotype: IGPS-60997 Hirose-mura, Miyagi-gun, Miyagi Prefecture Shirasawa Formation Late Miocene Acer momijiyamaense A. Takahashi et M. Suzuki, 1988 Bot. Mag. Tokyo, vol. 101, p. 474, figs. 1-7 Holotype: Dept. Biol., Coll. Lib. Arts, Kanazawa Univ., no. 22002 [TUSG], Isotype: TI and NSM-PP Momijiyama, Yubari City, Hokkaido Poronai Formation Oligocene [Late Eocene] Acer nomurai Okutsu, 1940 Saito Ho-on Kai Mus., Res. Bull., no. 19, p. 162, pl. 7, fig. 5 Holotype: IGPS-10987 Fukuoka, Nenoshiroishi-mura, Miyagi-gun, Miyagi Prefecture Fukuoka Formation [Shirasawa Formation] Late Miocene Acer oishii Tanai, 1970 Jour. Fac. Sci., Hokkaido Univ., Ser. 4, vol. 14, p. 490, pl. 14, fig. 1 Holotype: HUMP-25986 [NSM-PP] Okotsu pit, Harutori mine, Kushiro City, Hokkaido Harutori Formation Early Oligocene [late Middle Eocene] Acer palaeodiabolicum Endo, 1950 Short Pap. IGPS, no. 1, p. 12, pl. 3, fig. 3 Holotype: IGPS-60989 Kanchindo, Meisen-gun, Kankyo-hokudo, North Korea Kanchindo Formation Miocene [Middle Miocene] Acer palaeonippoicum Murai, 1969 Tech. Rep. Iwate Univ., vol. 4, p. 65, pl. 1, fig. 2; pl. 2, figs. 4, 6; pl. 5, fig. 4. Syntypes: IAGI-67009, 67010, 67011, 67012* Loc. 2, Yokokawame, Waga-cho, Waga-gun, Iwate Prefecture Hishinai Formation Late Middle Miocene Acer palaeoplatanoides Endo, 1950 Short Pap. IGPS, no. 1, p. 11, pl. 3, figs. 1, 9 Holotype: IGPS-60987

12 K.