14
/ / / ! "# $ %& (%)*% + $ , - ./ $ ( ($+ -’0 * 12 , 3 - $ ’2 450 36 78-2 9 & + : .- - + ;%0 < ) : / / : / / ( / = - =*%’ (% + -’# .$ <%& + > 0 - + ? 7 $ 50 @> A $ B <+ C D4# <+ E50 4F G0 ! (& 1% : - . ! I-+ $ +J C :# ’’ C "# .:0K (%)*% + $ G8 , - ./ 12 (% ( . . L6 + A F $ ) ! & M $ :0 N%& - O C . .0 $ # P + Q .& +- $ .& . .& ## R S T UVS MWWX T WXM U T V V WV T V 0 P P Q L6 % XW T X USX T XW W T SW T V XS T V 0 P P (%&# % MS T S UXV T M U T V S T V W T V 0 P P YQ Q% Z MU T UX MU T M T V S T V T V P 0 P % (%&0! - > . Q ! Q$ # Q+ Cyclotella Cocconeis Surirella P ! .& Chlorella Oocystis Tetraedron R[ . ! (%&Q# Q$ # Q+ Cyclotella Surirella P ! Q’ .& Mougeotia Chlorella Scenedesmus ! +- Closterium ! $ .& Prorocentrum ! .& Chroococcus Q R[ . $ ! Z $ # Q+ Cyclotella Surirella P! Q’ .& Mougeotia Chlorella +-’Q ! Closterium ! $ Q’ .& Ceratium ! QQ’ .& Chroococcus (%&QQ0! ! QQ$ # QQ+ Cocconeis Amphora P ! QQ’ .& Chlorella Tetraedron +-’QQ ! Closterium ! $ .& Ceratium ! .& Tolypothrix [ - R . \] + "Q# (Q (&Q^ T $ _Q‘0 Q . ## .Q0 R W / V T UV / V / / S / T S / V Q WW / V T S / V W / W / XV / T / (%&Q# W / V T SV / V / SM / X / T M / V $ Z M / V T / V U / W / V W / V T V / V (%&0! 7Q . Q’ %b $ "Q# C c%&Q$! Q (%&# . -0> 7 . 26] B d 1 # @> ! "# R $ %& (%)*% + $ C $ ./ . (9 $ e $ .Q/ Q$ I# 0 .;4F (# - . + f P - : (%)*% "# $ , - ./ . ! "#$%& ’()*% +,-& !’)./ 01*2 34,-& * : 2 56782 9 :& 8 !. 9 -( 9 : [email protected]

ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

Embed Size (px)

Citation preview

Page 1: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

������� ����� /������ /����� ��� /�������

��

���������� � ! "��# �$%& �'(�%)�*��% �+���$� ���, -� ./�$(�

(�$��+� ���-'0�*12 � , 3�'�-����$� ��'��2 �450 3�6� �78�-2 �

9'& +�� '� :� �'.��-�� -�+�� ';%0 < �

)����� ���� :��/�/������� ���� ���:�/�/����(

/='�-

=�*��%'�(�% �+������ -'��# .$��'<%& �+�> 0-��� �+� �'?� 7 $��50'� @> A'$������ B��� <+ C�D�4#�� �� <+ � E50

4F G��0 !� (�&�� ���1%�� �': -���� .!� I-+�$�+�J� C:#''C����� ������ �"��#� .:0�K '(�%)�*��% �+���$�� G8�� ��, -� ./�

��12� (�%��(�� ..���� ����L6� + A���� �� F $)!� ��&M�$:0 ��N%&'-� ��O�� C.����� .�0�� �$��#� ��P� ��+'�Q�� �.&'��+-

�$���'� � .&'����'. .& ##'R���STUVS �MWWXTWXM ��UTV�V�WVTV0 �� ��P� 'P �'���Q� L6� �� %�XW�T��X� �US�XT��XW �

�W�T����SWTV�XSTV0 �� ��P�'P �'�(�%&�# ��%��MSTS�� �U�XVT���M ��UTV�S�TV� ��WTV0 �� ��P� 'P �'Y�Q� ��Q%'� �Z

�MUTUX�M�UT��M���TV�S�TV����TV�� ��P� 0'P �'%(�%&0! �� -� ���> .��Q� �� � !� Q$��#� �Q+Cyclotella �Cocconeis �

Surirella ���P� !�'.&Chlorella �Oocystis �Tetraedron �� R��[ .� !� (�%&Q�# �� Q$��#� �Q+Cyclotella �Surirella ��P� !� �Q'.&

Mougeotia �Chlorella �Scenedesmus ��� !� �'�+-Closterium � !� �$���'.&Prorocentrum � !� �'����'.&Chroococcus ��Q R��[ .

�� ��$'�!� Z$��#� �Q+Cyclotella �Surirella���P�!� Q'.&Mougeotia �Chlorella �+-'�Q�� !�Closterium �� !�$��� Q' .&Ceratium !�

�'����QQ' .&Chroococcus(�%&QQ0! �� �!��QQ$��#� �QQ+Cocconeis �Amphora��P� !� �QQ' .&Chlorella �Tetraedron��+-'�QQ�� !�

Closterium�� !� $���'.&Ceratium � !� �'����'.&Tolypothrix [-��� R�� .\]�� �+�� "�Q�# '(���Q� �(�&Q^�T�$_��Q`��0 � Q�

. ##'.Q�0�� RW�/VTUV/V���/�–��/��S�/�T�S/V���Q� �� WW/VTS�/V�W�/�–�W/��XV/�T��/��(�%&Q�# �� W�/VTSV/V�

��/�–SM/��X�/�T�M/V�� �� $'�ZM�/VT��/V�U�/�–�W/V��W/VT�V/V�� (�%&0! �� 7Q� .Q'%b$"�Q�# Cc%&Q$! ��� ��Q�

(�%&�# .-0> 7��.2�6] B��� '� d�'1 �# �@> '! "��# � R$%& �'(�%)�*��% �+��$�� C$ �� ./�.(���9$�� e$.Q/� Q$ I�#�

0 .;4F (��# -������ .

���� �+�fP'-�:�'����� ������ �(�%)�*��% �� �"��# �$��, -� ./�

�.������� !"�#$%& '()*% ��+,-& !')�./ 01�*2 �34,-&

*:2 567829:& �8� !��.��9-��(�9':[email protected]

Page 2: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

������� ����� /������/����� ��� /�������

��

.0-;0

"6(9-��6(�� ;2� �< 6-��=>& ?���81?�(@�1 � �<A.BC ?�<

':6B= D��61 E@ ;����BF ?&�&a3*(8< )��(.�G#-H EG���

IH � 3GG�:6� � ��GG�J ��GG ?�GG<EGGK��� �6GG � L� � �GG<

E-�M�� �<'23* �1�?�G�> ���8G1 '�3G*F&�� �G)�./ �

3GG-�& .'GG2 &� '�&6GGC62 �GG(NF 3GG*-�N< EGGF �GG��� "&6GG�

"6(4-��6(�� �<O�GP2 � 3*G �1 ��Q RGB(S2 O�&�G � �G<

3G**F 3 � ."6G(4-��6(�� � '4�T6G:6F& 56G%& U�G=&�1 �G<

V< "�8-& ��$(=& �1 W�.��& � L�*K �G�N<& 'G)�./ 01�G*2 >&

3-�& '�X�1 .G1 '-6G(4-��6(�� �6G� �8G�> �>��G2&E"&6G*Y

!?>�G=�Z�F !'(8G�> ?�G<6F !' ���� "��[1H � \& ?&�Z

?T6:6*4� O�P2 L�6- ?�< EG-�M E�$]G� � �8G�> IH ?�G<

E1 I^�GC "&6G*Y ��$(G=& '(8G�> �M6G= � !'(8G�> ?�G<

'2 3-6 )_!�_���(.

�`2 >& ?���8GGGG1 � a6]GGGGM � '4�T6GGGG:6F& ��GGGG):

"6(4-��6(�� !�< '-�42 � '-�2> ��Y6*� �=& EbJ�2 ;1�Q .L�1

��"6(4-��6( &�G � �<��G� I�GS(-& � �G#-H '�3G-> � 3G � O

���$� L�)2 � cS,2 '`�P2 ?�<�6(F�� ;NP� 6GC� '��G<

�&.EG-6� �6dJ \3Y � �6dJ !�G#-H e*F&�G� ?�G<6+:& !�G<

E)2�C "61 "6��-6���"6(4-��6( G(2 �G#-H ��Y�N(C& � �<f>& �g

�>&3G-& ;G1�Q EGF �=& '`�P2 ?�<�6(F�� G2 ?�G��'�G2& !3G �1

��"6(4-��6( � �P� ��&6N< �<fL�G1 ;G1�h(2 ��gH � ��/�.��& ��g

E]S,2 ?�G<�(2&��� ���G= �G1 L�)2 � �3 E(M�* '`�P2 ?�<

�>&3-& ;1�Q��Z 3-�& �&�Q ��b(-& ;1�Q��Z � ?��� .!LG�& �G1 ���Y

!'GN�BQ& O�&�G � 6GM e*F&�G� � �G=& LG4N2 E-6� A�

i4Y 3-&6(1 '(M�* �8�> � '����&�jC EF 3 �1 E( & 'BN):&

�Gk�& O�&�G EG1 'B��N� � �=& �6N- E1�k� &� "H E( �� �

3 �1 E( &3- '-6*F �3 .1EEG-6� 'BF �6/ � ?�G<�'-6G(4-��6(

E.*C �1 ���b- ?&�1 �)2 ?�< 0��GQ� EBNC >& �8�> O�P2 >& '*

"3G \�� \�Q�& � ��2H !'�6:H !"6�=�4$���6� !'4��&����3�<

�8�> ��4,2 ����8G1 �3G2 >&� �&�G��j� l�P: E1 '`�P2

3*(8< 3�$2)��.(��"6(4-��6( m`= � �<�X ��G�& E':�*NG�"6

��EK� �)� � �<�Eh./ � R3*1?��G���� m`= l�P: E1 EK�'

(trophic state) :6���:�& 3G�N<& EG�3G-�& �.G1E ���G.Y��G+

F��GG�EGG-6� n&?��"6GG(4-��6( �/ >& �GG<GG�:&6GG� o'B]GG� '

(seasonal succession)> p6GGGG*� !GGGG�(8'(biodiversity) !

E-6� �<?1�3-&6��6��4(bioindicator species)cM� � !�<?

>GG�(8'(bioindices)GG���� m`GG= �GG1 'GG�� A�.��GG2 EGGK� O

'2 6 )�q.("6(4-��6(�� �6dJ '�3�r�� E1 EC6� �1 � �<

���G� �G1 �&6GC62 LG�& ���G$(2 OG1&�� � O�P2 �< ?�G<

���� �M�* !�+� '(8�> ���81 '-6(4-��6(�� RB(S2 ?�<

�& ��N<&.

T6:6F& ��):�`2�4'��"6(4-��6( EG1 �< � ?�6G2 ��6G%

IX�� EK��� � �< 3 \�k-& "&��& ?�< �=& �)�!�!�!s��.(�1

& E1 EC6��L� EF�"6(4-��6( &�& �<?���� �<?$B(S2 '3*(8G<

� EFfg�Y6*(2 �&�':6� �1 �:�& 3��j� ��/�.��& !E�E&?� tG-� �

%6]M�F ���$'(= &�: 3-�& IH �1�'E1�6+:& A?nG=�*2

N)� ;1�Q ��&�1 V?� o/�*2 �EK� &?6GC62 ��GY�/& u3-& �1

&6 �=& �.�� EG):�`2 ����Gv &�G: G�E&?& � G�a6]GM L

6:�& � "�*rN<�61 3<&6M �.��(=&�?1 ��):�`2 G��3G "�

-&�&�� !e*F&�� E):�`2'F�� ��E-6� n&?� E)2�C �"6(4-��6( �<

���2& �*J 3= EK�?���v ?2'3 �1 .&�� L�V#2 >& EK� L���

V(8�=6F& EF �=& "�#$%& "�(=& � '1H ?�< � �q?�(26GB�F

\��N= "�(=�# )��q "�#$G%& w�G I6G*C ?�(26GB�@ (��

x�Y '��G��&�jC°���'��� ':�NG 56G/ '��G��&�jC °���

's{'Q� �=& 0Q&�);4 �(.� L�& �EK�3*-�2 G�IX�G� A

��GG�J !�C�GG#2 "��3GG-�� ?&�GG1 '4�T6GG:6F& �GG�N<& � eGGJ�

�& "�G�<�2 a6]GM E1 ?[1H �&6C62 .>&�Gb- "�G�<�2 "6G��

':6GF ;2�G '261 '<�2 E-6�)Alburnus sp.([G�� '<�G2 ���G= !

iGB�)Capoeta damascina ( U6�-�G�H �)Aphanius isfahanensis(

'GG��/ U&��GGF '<�GG2 EGG-6� � �)Carassius auratus (EGGF

5[GGGQ '<�GGG2 � '26GGG1 &�GGG�& OGGG=6� "�GGGNF L�GGG+-� ?XH

)Oncorhynchus mykiss (1 ��� O=6� EF'G��)2 EK��� E

�3 3-&)�.(

Page 3: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

����� %&$! "��# � ����� �(�%)�*��%'� (�$� ���, -� ./�$�� ��+

�{

& E1 EC6� �1�LE2�-�1 EF �<?IH �G1 ���Gb- �G<?G ����G(#1 L

�=�* U�=&�1 �=&'����� �<?26-68F�� RB(S2 �4'�� 3 �1

[<�E*�<?E2�-�1 �<?���b- '&�1 ?32���& �G�IH Le<�GF �G<

�31� .=��1 &�:'-&�&�� !e*F&�� 'p6*� ���"6(4-��6( �<��EK�

�1��F ��Y�/& 5�= RB(S2 56]� � �*J 3=?T6:6F& �4'�

1�T6:6�4'&�1 &� ?� ���b- � '4�T6:6F& "&6� L��)� ����3G2

� L�& '�H�� � EK��EK� 6GN- 3G<&6M V<&�� E1�,2 ?�< .�G�&

& � EGGKGG� GG=��1 e<�|GG� L'j�GG����(M�GG= �&�?EGG)2�C

��"6(4-��6( � �< )� �G� G���� m`G= L'�EGK�� G=��1 �6G2 '

�& "�N� �2 �2& �����&�Q�32 EF V����3G2 >&� ���Gb- � �

&�� L�2 EK�'� EG)2�C ��(M�G= U�G=&�1 3G-&6��-6G(4-��6('!

F�$�IH ��=��1 'j� 3-�� ��� �&�'� � '6 \�k-& .

h�� � ���0 �+

� � .;i�0 �:0$��N%& �+�.���� ����

C ��)Q62� '����&�j�;4G � �*J 3= EK���3G cS,G2

�GG=& .�GG#C oGG�hP� LGG�& � �GGb- �6GG2 }&3GG<& 'GG=��1

E-6N- "�(81�� >& 5�= A� �32 E1 ?�&�1��_��G� ��G#1��_{

L)��.�� .b;� $��N%&$� 7':8�0 � ��, ./�.���� ��+F�Q�0 �� ����j�����)S1, S2 ,S3

0i�.;��S4 �S5 .;i�0��S6 .;i�0�(.

º��,´��,"sqN

º��,´s�,"��E

º��,´��,"��N

º��,´s_,"��E

º��,´�s,"{N

º��,´s{,"��E

1SS2 S3

S6

S5

S4

Page 4: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

������� ����� /������/����� ��� /�������

�_

� AG�T�� Eh`*2 >& ;]� �< O=&�& ��\�Gk-& �G*J 3G= EGK�

3��� .E):�`2 L�& � ���$� E1 EC6� �1 �<?L�+-��2 � eJ��

� oNY T6G:6��2 ���G$� � IX�G� E1 IH ?��� !EK���4'�

;4 EG-6N- ?&�G1 EGh`*2 EG= "H �)G=� � IX�� � ?�&�G1

��3GG EGG(��� �GGb- � 3GG= EGGK�);9GG �.(EGGh`*2�);GGP2

��+(8�& ?�G<�!���(L�G1 oGNY OG=6(2 ?&�&�q��q!�G(2

���3NY �(81 !IH ?��� "�31 EG-& �G1 � '+*= !�G � ?3G*1

Eh`*2�)��+(8�& ;P2 ?�<s��(L�1 oNY O=6(2����!�(2

IH ?��� ?&�&)�3GGJ�qIX�GG� IH ?��� >& 3GG%� (!

E-& �1 �(81 E=�2 � O=6(2 ?3*1 � ���* ?[1H "�<��� ?&�& !?&

E/6Z Eh`*2 !���)��+(8�& ;P2�(L�1 oNY O=6(2�q���(2

���3NY �(81 �1 !���8G1 'G:H &62 ?&�& !'=� IH ?��� ?&�&

)�3J�q� IH ?��� >& 3%� �EK�(?[G1H "�G<��� ?&�& !

E/6Z �=& IH � ���* � �� .1 ���$� ;4 �< ��o/�G*2 L

�h2 ��& E1 E8���+(8 �<?81 o/�*2 "�� �6G1 EbJ�2 ;1�Q �� .

� �J�82�EK�{qq �G�� m`G= >& p�$��& !��(4< ��qq !�G(2

IH oNY E-X�= L�+-��2�qEh`*2 E-X�= '�3-��1 O=6(2 � �(2

�_q 'B�2 �(2 '23 �1 .L���� O=6(2 &6G< ?�G2 L���X�1 � L���

E1 LN#1 � n����)�/�'(-�G= EGC� &�G� (&�G2 � �)�/��

'(-�= EC� &�G� ('G2 3G �1 .� �G= V�GBQ& ?&�& EGh`*2 LG�&

1 �=& '-�(8<6@E?�6/ ;]G� >& 'GN�- � EGK��� m`G= EG@

82> '2 � >& �3� 6� "�( 3 �1 .eGS1 EGK��� oGNY VGF ?�G<

1Ei*C >& IH � L1 "�<��� EB�=� ?�G<Phragmites !Typha !

Juncus �Cyperus'2 5�j & ':�J � 6 EG/6Z "�<��� EF��

i*C >& ?�<Potamogeton !Ceratophyllum !Myriophyllum

�Polygonum3-�& 6C� �#2 �� �,#.��& >& .

�.����!�-�� � @> ���� $�'�'��=$Z'� ��+��%f�� �'`

EGG-6N- VGG#2 L�GG�)� �GG#C IH >& ?�&�GG1 ?�GG<�(2&��� L��GG�

�Y�GG= >& IH '���N�GG 69�[����GG� ��>& ��$(GG=& �GG1 m.GG%

E-6N- IH �&�1Van Dorn m`= >& )>& �(NF��(2 ("��2 !)��G�

��(2 (oNY �){���q�(2 (Eh`*2 E= �);4G �(EGK��� >&

�G��� \�Gk-& �*J 3= .;2�G IH '���N�G 69�[�� ?�G<�(2&���

!�2pH !oNY � 56BP2 "|�8@& �G1 EGh`*2 �G< ;GP2 � ����

�6GG�C �*GG=�2 !?& pH �GG(2)5�GG(�k�Schottgerate 53GG2 !

������!"�N:H �M�=(�G(2 "|�8G@& !)53G2Paqualab ELE !

"�N:H �M�= (�>&3-& ',4= EP$% �3 ?���.EG:�h2 LG�& �

;]G� �G< ?&�G1 oGNY EG= >& �Gb- �6G2 ?�G<�(2&��� L�+-��2

3G ��&[� .EG-6N- oGNY >& IH ?�G< o/�G*2 � ���G$(2 ?�G<

E= G1 � 3G ;h(*2 ��+,��2>H E1 E-��E� �&�G(�- EG-��&3C �6G/

�>&3-& �#-H ��$8� 3 ?��� .�&�(�- 1E�+1�GS(-& ��G(9:& EB�=�

'-6G�)Ion Selective Electrode !53G2 JEAN WAY 3310 !

"�N:H �M�=(t-� ��� E1 56BP2 ��$8� !��$(G=& �G1 'k*=

�(26(���(9�GG=& >&)53GG2JEAN WAY6400"�GGN:H �M�GG= !(

�>&3-& 3 ?���)�s(.� ��$8G� � �&�(�- ���(- L�+-��2 �EGK�

�=& �3 ��&[� E:�h2 L�& � ;]� �< �.

.���� ����� C'':# � $����� ������'(�%=�*��%�+

E-6N- "6(4-��6(�� >& ?�&�1 0GNC �1 �< oGNY >& IH ?��H ?�G<

L�1q��\�Gk-& �G#-H "6GN- W6GBS2 >& i� '��F "&[�2 E1 �(2

3 ."6G(4-��6(�� ��.�� �#C EG-6N- EG1 �G< IH ?�G< 56GBP2

L�6�H 56�6:)�q'B�2 �< ?&>& E1 �(�:�qq 'GB�2 EG-6N- �G(�: (

=�* � 3 E��v& i*C '�� ?�6G- �69=��9�2 >& ��$(=& �1 �<

'��GG=�* ?�<3GG�B@ �)�q!�����(�GG��� \�GGk-& .���NGG

��"6(4-��6( �3G= \X >& ��$(G=& �G1 �G<����NG � �G�&� A

\�� �<?E( � &?��"6(4-��6( �>&3-& U�=&�1 �< ���?E( � 56/

2 >& ��$(=& �1�N,K �(2��4'3 \�k-& )�q.(

\]�� .4��50 �+�"��#

EGGG-6� p6GGG*� EGGG):�`2 LGGG�& � 'BGGG%& E(GGG= EGGG= � ?&

E-6� ?�*Z cM� ?&)species richness ( cM�G U�G=&�1

R:�GG���2)Margalef index(�GG�.:�Z !/EGG-6� '(M&6GG*4� ?&

)species evenness/dominance ( = cM�G U�=&�1�"68G�N

)Simpson index (F�� ���*Z n?.:�Z �G�EG-6� �&?U�G=&�1

Page 5: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

����� %&$! "��# � ����� �(�%)�*��%'� (�$� ���, -� ./�$�� ��+

��

��-K�.�� �� _P%m0 ��6� �� @> 1'� ��+%0���� $��, ./�)C'N��'0±���-��%�� ��i] .(C'N��'0 L8�-, ����� .f _$�� + �� �+

oi� �� ���0> p� !� -�%&+ q%b0 I, r$VS/V �:0 I*%]� C=��� (�0!> � -2�� -���-� ����.

"�(82> [���� "�(81�� ��#1 �(2&���

�s/q±�s/� d s�/q±��/�� c s{/q±��/�q a ��/q±{�/�s b (ºC)IH ?�2

q�/q±{�/� a q_/q±�s/{ b ��/q±s�/{ b ��/q±q�/_ b (mg/L)56BP2 "|�8F&

�/�±�/�� c s/�±�/��_ a {/�±�/�q� b �/�±�/�_ c (cm) ',4= ���� oNY

q�/q±��/_ a q�/q±�s/_ b q�/q±�s/_ a q�/q±q{/_ c pH

q�/q±�{/� a q�/q±_�/� bc �_/q±��/� b q{/q±�_/� c (mg/L)�&�(�-

qq/q±q�/q c q�/q±�{/q b ��/q±�q/� a q�/q±q�/q c (mg/L)��$8�

cM�GG "6-�GG ����GG*)Shannon-Wiener index (� �6GG2

GG=��1 � E.GG=�P2'�GG��� �&�GGQ .cM�GG �GG<?=�� "68GG�N

"6-� ���\�G- >& ��$(=& �1 �* �&[G�&Ecological Methodology

)��(U�=&�1 R:����2 cM� �R:�G���2 ��Gh�hP� )�q!��

���(�� E.=�P2�3.

���>'�+Z����0>

�& cM�G � IH '$�F ?�< �G<?[�:�G-H >& ��$(G=& �G1 p6G*�

E��/ A� i-���&�)One – way ANOVA (�62 ?��2H E�[k�

L�+-�G�2 � !���� �&�Q �6G2 LG4-& "6G2>H >& ��$(G=& �G1 �G<

�G��� �&�Q E8��h2)��.(:�-H \�GN� �'G*)2 m`G= � �G<[ ?�&

q�/q\�- >& ��$(=& �1 �&[G�&)SPSS; version 11.5 ( 3G \�Gk-&

)�q.(

s$�%�

@> $�'�'��)$Z'� ��+��%���

��]S,2 IH '���N� 64�[�� � �� EK� 5�3C��3G E�&�&

�=& .?�G2 'B]G� L�+-��2 EG1 56GBP2 "|�8G@& � IHnG����

C◦{�/�s�q�/_'B�2 !��G#1 � �(�: �1 \��C◦��/�q�s�/{

'B�2"�(81�� � �(�: �1 \��!C◦�� /����s/{'B�2 �(�: �1 \��

�[���� �C◦�s/��{�/�'B�2 6G1 "�(8G2> � �G(�: �G1 \��.

]� L�+-��2 'BpH E1 ',4= EP$% ���� oNY �n���� q{/_

��/�_'(-�= !��#1 � �(2 �s/_��/�q�'(-�= !"�(8G1�� � �(2

�s/_��/��_'(-�= � [���� � �(2 ��/_��/��'(-�G= � �G(2

6G1 "�(8G2> . ��$8G� � �&�G(�- 'B]G� L�+-�G�2 [G�- 56GBP2

E1 n�����_/��q�/q'B�2 ��G#1 � �G(�: �1 \�� !��/���q/�

'B�2 !"�(81�� � �(�: �1 \��_�/���{/q'B�2 � �G(�: �G1 \��

� [�����{/��q�/q'B�2 G1 "�(82> � �(�: �1 \��E3G2H �G=

)5�3C�(.

�#'.��` R������� � �'(�%)�*��% �+

F���E-6� n&?G-&�&�� �'��"6G(4-��6( �G<?& � G���+(8 �G<?

E-6N- �3 ?�&�1 � RB(S2 56]� � �*J 3= EK��� 5�&3GC�

����=& �3 E�&�& .��G#1 ;]G� � i*GC ?�G< >& '-6G(4-��6(��

\6��GG� �GG<)Bacillariophyceae ( E8GG����BF �)Chlorophyceae(

)NCGG� 4,GG� &� nGG:�Z ��:�GGJ � & ;'NGG= >& EGGF��<3

)Desmidaceae ( i*GGGC �GGG#*�Cosmarium >& ���6*GGG� E8

)Dinophyceae ( i*GGCCeratium�GG=�* '��� GG�3.EGG*2&

-&�&��'��"6(4-��6( & � ��#1 � �<���+(8 �G<?&�G1 RGB(S2 ?

�\6�� �<�����sq� ���BF !�E8�{{_�{_�N= !��<3�s�q�

��6*�E8{q�q2 � 56B= �B':�1 �(E3G2H �=)5�3GC�(.�

��#1 G�\6�� �G<?i*GC nG:�Z �G<?Cyclotella !Cocconeis �

SurirellaGG-&�&�� EGG*2& EGGF 6GG1 'EGG1 �GG#-H ���GG� n�{���{�!

{_���q� �����_�2 � 56BGG= �GGB':�& � �GG(GG���+(8 �GG<?

6GG1 RGGB(S2 .GG1E#1�,GG2 �6GG/'���BF � GG�i*GGC !E8 �GG<?

Chlorella !Oocystis �Tetraedron -&�&�� E*2& � 61 n:�Z '

Page 6: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

������� ����� /������/����� ��� /�������

{q

��-K�.�# '.��` R��0 �'N��'C)±�i] ��%�� ���-� (����� )0 �� ��P�'P �'%(�'(�%)�*��% �+�./�$�� ��,

� �� ��� L6� ��$��N%& �+�_P%m0

��+(8�& E-6N- ?�< ?�&�1

� � s � � �i*C �

s/{q±�/�{ s/s±s/� q q _/��±�/�� q Navicula �/{__±�/�q� �/���±�/�� {/s�s±�/�s� s/�{{±�/�_� �/�_�±�/�� �q�±s/ss Cocconeis �/s�±�/�� q q q q q Amphora

q q �/��±�/�� q q q Encyonema �{�±�/��_ �/�{�±�/�s s{s±q s/{��±_/{� �/��q±�_ �/���±�/�� Cyclotella

�/���±�/��_ �/�_s±�/�� �/���±s/�� _/��{±_/{� �/���±{/�� �/_�±�/�_ Surirella �/���s±s/�s� _/���±�/�q� {/��_�±�/��s {/���_±�/��� �qs�±_/��� �/sqs±�q� L�+-��2

\6���

�<

_/���±_/�� �/��±�/�s s/��_±�s {/��_±_/�q� s/���±�/�q{ �/�q±q Scenedesmus

q q �/�±�/�� q q �/�±s/_ Pediastrum �/����±�/ss� �/�_s±_/_� �/���q±s/��� s/�_q�±�/{_� �/�qqq±�/��� �/���{±s� Chlorella �/�_�±�/�� �/�_�±�/�� �/��±�/�� ��_±�/�s �/�s±�/__ W/��±X/�M Tetraedron �/�_±�/�{ q q �/��±�/s_ q V Mougeotia _/���±_/�� �/{s±�/�� �/�s_±�/�� {/�{s±��� �/���±�_ {/�q�±�/�q� Oocystis

�/����±�/��� �/{_�±�/��s �/�_{�±_/�{{ �/�{{{±�/

���� _/��_�±�/�_� ��sq±s/�{q L�+-��2

E8����BF

q q q q q {/��±_/�� Cosmarium

q q q q q {/��±_/�� L�+-��2 �<3�N=

s/{q±q q q q q q Ceratium

s/{q±q q q q q q L�+-��2 E8��6-��=

EGG1 �GG#-H ���GG� n�_q���_�!�{��{����_��s� 56BGG=

2�B':�61 �()5�3C�(.

F�� "�(81�� ;]� ��E-6� n&?-&�&�� �'��"6G(4-��6( �G<

&� ��81 O�2 �� � EF � & �&�Q n=�* 5�3C��3G EG�&�&

�=& .& ����+(8 �<?E-6N- �&�1?-&�&�� '�\6�� ���BF !�<�!E8

N=� !�<3��6*�= � E8��6-� G� E8(Cyanophyceae) EG1 ��� G� n

�_{����_�!s��_���_{ !�{����!��{�q�_��q56BGG=

'GGB�2 � 6GG1 �GG(�: .i*GGC \6��GG� nGG:�Z ?�GG< �GG<Cyclotella �

Surirella E1 �#-H '-&�&�� E*2& EF 61 nG�������q�{qq ��_��

��� 'GB�2 � 56B= E8G����BF >& !�G(�:Mougeotia!Chlorella �

Scenedesmus E1����-&�&�� E*2& n'�q�{���!��q���q��

��{�����2 � 56B= �GB':�NG= >& !�G(��<3Closterium �G1

-&�&��'�s_�q2 � 56B= �B':��( >& !��6*�E8Prorocentrum

GG-&�&�� �GG1'��s�q2 � 56BGG= �GGB':�= >& � �GG(��6-� GG� E8

Chroococcus -&�&�� E*2& �1 '_��q2 � 56BG= �GB':�6G1 �G(

)5�3C�.(

�(-�F�� ��-&�&�� � n'��"6G(4-��6( �G� ;]G� � �G<��� [

5�3Cs�h2 � EGF �=& �3 E�&�& G�E8�G1 �G� "�(8G1�� ;]G�

Page 7: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

����� %&$! "��# � ����� �(�%)�*��%'� (�$� ���, -� ./�$�� ��+

{�

��-K�.�# '.��` R��0 �'N��'C)±�i] ���-��%�� �(����� )0 �� ��P�'P �'%(�'(�%)�*��% �+���$��, ./�

� �� (�%&�# L6� ��$��N%& �+�_P%m0

��+(8�& E-6N- ?�< ?�&�1

� � s � � �i*C �

�/��_±�/�� �/{�±�/s_ �/���±�q �/�{±q s/_s±�/s� {/�_±�/�q Navicula _/���±_/�� s/��±�/�� �/{{±_/�{ q/_�±�/�� �/���±�/�q� �/{s±s/s� Cocconeis �/��±�/s� �/��±�/�� q �/�{±q �/_s±{/�q �/{s±s/s� Amphora _/���±�/�s s/��±�/�� s/��±�/�_ q q q Coscinodiscus �/���±_/�� �/{�±�_ s/���±�q �/s�±�/�� {/�_�±�/_� �/���±�/�� Nitzschia

q q s/��±�/�_ �/�±�/�� q �/��±�/�� Fragilaria �/�s±s/�{ �/��±�/�� _/�q±�/�_ �/�_±�/�� �/�q±q q Encyonema s/_�±�/�q �/��±q {/��±�/�_ �/�{±q �/�q±q {/�s±�/�� Gomphonema

q q _/�q±�/�_ q q q Asterionella _/���q±_{� �/{qq±�/{q �/���±�/_� s/���±�/��_ q/���q±�/�s{ �/{q�±_/s�_ Cyclotella {/s�±�/�� �/��±�/�� q q �/�{±�/s� {/�s±�/�� Cymbella �/�s±s/�{ �/��±�/�� q �/�±�/�� �/���±�/s� �{±q Synedra {/�{q±�/��� s/�{{±�{� �/�_�±s/��� �/��_±�/{� �/���±�/�q� �/��q±�/��� Surirella �/��_{±_/��_� �/��_�±�/s�� _/��q�±�/�_� q/��{�±�/��� �/�_{�±{/�_� �/�s_�±�/_�� L�+-��2

�\6��

�<

�/�s�±�/��_ {/�{�±{/�s �/���±�q �/��_±�/�� �/�{�±�/s� �/�s�±�/�s Scenedesmus �/�_±s/�{ �/��±�/�� q q �/�q±q �/��±�/�� Pediastrum �/�q��±s/�q� s/��q�±�q�s {/�__�±_/�_� �/�{s�±�/�s� �/��q�±�/�q� {/����±�/���� Chlorella

q q s/��±�/�_ q q V Coelastrum

s/_�±�/�� s/��±s/�� q �/�±�/�� �/�q±q {/�s±�/�� Tetraedron {/_��±�/{� ��{±�/��� �_�±�/�q� �/��±�/{_ _/sqs±q/�� _/�q��±�/��� Mougeotia �/�_±s/�{ s/��±�/�� �/s�±{/�� �/�_±�/�� �/�q±q �/{s±�/�� Oocystis

s/���q±s/_sq �/�_�{±�/��{� �/�{��±�/s�� �/��_�±�/�_� �/���{±�/�q� �/s��{±�/��q� L�+-��2 E8����BF

q �/��±�/�� q q �/�q±q �/��±�/�� Desmidium �/�s±s/�{ q s/��±�/�_ �/�_±�/�� q �/��±�/�� Cosmarium

q q _/��_±�{q �/{�±�/_� q �/�s_±s/�_� Closterium �/�s±s/�{ �/��±�/�� �/��s±�/�__ �/��±�/�qq �/�q±q _/�{�±�/��� L�+-��2

�<3�N=

{/s�±q s/��±�/�� q �/�±�/�� q q Ceratium _/���±{� q q q q �/��±�/�� Prorocentrum �/���±{� s/��±�/�� q �/�±�/�� q �/��±�/�� L�+-��2 E8

��6*�

q �/��±�/�� q q q q Agmenellum

q �/��±�/�� q �/�±�/�� q q Aphanizomenon

q s/��±�/�� _/�q±�/�_ �/�±�/�� q {/�s±�/�� Anabaena

s/_�±�/{� �/��±�/�� q q q �{±q Chroococcus

q q q q q �/��±�/�� Tolypothrix

s/_�±�/{� ��±{/�s _/�q±�/�_ �/�_±�/�� q �/{s±�/�q L�+-��2

E8��6-��=

Page 8: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

������� ����� /������/����� ��� /�������

{�

��-KU.�# '.��` R��0 �'N��'C)±�i] ���-��%�� �(����� )0 �� ��P�'P �'%(�'(�%)�*��% �+���$, ./� ��

�� L6� ��$'� �� Z$��N%& �+�_P%m0

��+(8�& E-6N- ?�< ?�&�1

� � s � � �i*C �

�/{�±�/�s �/��±_/s� �/{{±�/�_ q {/��±{/�q �/��±�/�� Navicula �/�_±�/�s q q �/�_±�/�� q q Cocconeis �/�s±s/�{ q �/s�±{/�� q {/��±�/s� q Amphora �/�s±s/�{ q q �/�±�/�� {/��±�/s� q Coscinodiscus {/s�±�/�q s/��±�/�� �/��±q q q �/��±�/�� Nitzschia

q q _/�q±�/�_ q q q Fragilaria q �/{�±�_ {/��±_/�{ �/�qq±�/�� q q Encyonema q q q �/�_±�/�� �/��±{/�q q Gomphonema q q q q �/��±{/�q q Asterionella

�/���±{� �/{��±�/��_ _/{�s±�/�q� s/���±_/�� s/���±��� {/s��±�/�q Cyclotella q s/��±s/�� _/�q±_/�{ q �/��±{/�q q Cymbella

_/���±�/s� s/���±s/�_ �/�qq±�/_� �/��{±�/�{ �/��_±�/�ss �/���±�/��� Surirella

{/_��±_/��� �/��q�±��� �/���s±{/sq� �/_��±_/��{ �/����±�/s�_ �/���±�/��� L�+-��2

\6���

�<

_/���±�/s� �/��_±�q� �/�q{±�/�_ �/��±�/�� {/��_±{/�s �/�s_±s/�s Scenedesmus q �/��±�/�� s/��±�/�_ q q q Pediastrum

�/��s�±�/���s �/s�qq±��q �/��{�±�/��� {/�_��±�/�{ ���q±{/��s� �/���q±�/�{� Chlorella q q q �/�±�/�� q V Coelastrum

�/�_±s/�{ q �/{{±�/�_ q {/��±{/�q {/�_±�/��q Tetraedron {/��_±{/��� _/��q±�/{q s/�q_±_/��� �/��{±{/s_ �/��q±{/�s {/��q±s/�_� Mougeotia {/�{q±{/��� s/��±�/�� s/��±�/�_ q {/��±{/�q �{±�/�� Oocystis

�/���_±_/��_� s�_q±�/��� �/�q�{±{/��� �/��s�±�/��_ �/����±�/�{qq �/����±{�q L�+-��2 E8����BF

q q q �/�±�/�� q q Cosmarium �/�s±s/�{ q q q q q Staurastrum

�/�s±s/�{ q q �/�±�/�� q q L�+-��2 �<3�N=

q q q q {/��±�/s� q Ceratium q q q �/�±�/�� �/��±{/�q q Pyrodinium

q q q �/�±�/�� �/�q±�� q L�+-��2 E8��6*�

q q q �/�±�/�� q q Aphanizomenon

q q q �/�{±�/�� q q Chroococcus

q q q �/��±�/�q q q L�+-��2 E8��6-��=

Page 9: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

����� %&$! "��# � ����� �(�%)�*��%'� (�$� ���, -� ./�$�� ��+

{�

��-KS.�# '.��` R��0 �'N��'C)±�i] ���-��%�� �(����� )0 �� ��P�'P �'%(�'(�%)�*��% �+���$��, ./�

� �� (�%&0! L6� ��$��N%& +��_P%m0

&���+(8 �<?E-6N- �&�1?

� � s � � �i*C �

q ��±�/�q q �/���±�/��� q q Cocconeis q �/�{±q q q q s/��±s/{s Amphora q q q q �/�s±�/�� q Nitzschia q ��±�/�q q q q q Gomphonema q q q q �/�s±�/�� q Cyclotella q �/��±�/�� q q q q Cymbella q �qq±�/{� q �/���±�/��� �/s_±�� s/��±s/{s 2�+-��L

�\6��

�<

�/�q±�� q q q q q Scenedesmus �/��s±�� �/_{±�/�� ��q±�/�� {/�_q±�/�s �/�ss±q �/��q±�/��{ Chlorella

q s/s��±�/{q �/�±_ q �/���±_/��s �/��±{/��s Tetraedron �/sqs±��s �/s��±s/_� �/���±�/�s {/�_q±�/�s _/sq�±_/��s �/��s±�/�_� 2�+-��L

���BF

�E8

q �/��±�/�� q q q q Ceratium

q �/��±�/�� q q q q 2�+-��L

��6*

�E8

q q �/s�±_ q q q Anabaena q q �/_�±�/�� q q q Tolypothrix q q �/���±�/�s q q q 2�+-��L

=��6-�

�E8

�3J?�=& �3 ���3P2 .E*2& -&�&�� '�\6�� ���BF !�<�!E8

NGG=� !�<3��6*GG�= � E8��6-�GG�EGG1 E8���GG� n��������!

s�_q����� !�s�q!���q� !�{�q2 � 56BGG= �GGB':�� �GG(

&���+(8 �<?1 RB(S2 E3G2H �= .��"6G(4-��6( �G<?>& nG:�Z

�����;2� \6��Cyclotella �Surirella E1����G-&�&�� �1 n'

����s�{��������2 � 56B= �GB':����BF !�G( G� ;2�G E8

Mougeotia �Chlorella EGG1 ���GG� n������_ �s�q������

2 � 56B=�B':� >& !�(��6*�i*C E8Ceratium -&�&�� '�s�q

2 � 56B=�B':�= >& � �(��6-� G� i*GC E8Chroococcus EG*2&

GG-&�&��'�{�q2 � 56BGG= �GGB':�& � �GG(GG���+(8 �GG<?RGGB(S2

E-6N- �&�1?� & &� .

GG-&�&�� "�(8GG2> �'GG�\6�� ���BF !�GG<GG� !E8��6*GG� � E8

=��6-�GG�EGG1 E8���GG� n��s�s_!��s����!���q!���q�

����q2� 56B= �B':�G1 �G(E3G2H �G= .G�\6�� �G<?i*GC

Cocconeis �Amphora E1����G-&�&�� �G1 n'��s�q����q

2 � 56B=�B':����BF !�( G� �G E8;2Chlorella �Tetraedron

E1 ����n����__�s���q2 � 56BG= �GB':� >& !�G(��6* G� E8

i*GGCCeratiumGG-&�&�� '���q2 � 56BGG= �GGB':�>& � !�GG(

=��6-��i*C E8Tolypothrix -&�&�� E*2& �1 '_��q� 56BG=

2�B':�& � �(���+(8 �G<?EG-6N- RGB(S2 �&�G1?3G-61 nG:�Z

)5�3C�(.

]��\�+�"��#

cM�GG �GG<?� p6GG*� �"6GG(4-��6( � RGGB(S2 56]GG� � �GG<

Page 10: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

������� ����� /������/����� ��� /�������

{s

&���+(8 �<?E-6N- �&�1?� �3 �� �*J EK� ;4 ��3 E�&�&

�=& .�(-�cM� ��#1 ;]� � EF & "�,- ��<?=�� "68�N

"6-�GG ���EGG1 �GG* ���GG� EGG*2& n{�/q�sq/q���/�–��/�

1E1 EF 32H �=��(,�& � L���+(8��(NF ��& � "H LG���+(8

�61 ."H E*2& ��#1 � R:����2 cM� ��/����/qEGF 6G1

1��(,��(NF � L�E1 "H L����& � n���+(8���61 .;]G� �

cM� "�(81�� �<?=�"6-� � "68�N���EG1 �G* ��� G� EG*2& n

{{/q���/q�{�/�–�{/�GG1 E1 EGGF 3GG2H �GG=GG��(,�� L

&GG� ��+(8��GG(NF ��L& � "H GG� ��+(8�)= U�GG=&�1�"68GG�N (

&����+(8�)"6-� U�=&�1����*(6G1 .� R:�G���2 cM�G

&�"H E*2& ;]� L_q/����/�1 EGF 61 G��(,��G(NF � L�"H L

E1 ����& � n���+(8��s6G1 .�G� ;]G� ���cM�G [�G<?

=�"6-�GG � "68GG�N���EGG1 �GG* ���GG� EGG*2& n{�/q��q/q�

��/�–��/�1E1 EF 32H �=��(,��G(NF � L�EG1 "H L��� G� n

& �GG� ��+(8s��6GG1 .- R:�GG���2 cM�GG GG� "H EGG*2& [

_�/����/q1 EF 61 ��(,��(NF � L�E1 "H L����& � nG���+(8

s��61 ."�(82> ;]� � cM� �<?=�"6-�G � "68G�N�

��E1 �* ����E*2& n��/q���/q�s�/�–�{/q1E3G2H �G=

EF 1G��(,�& � LG���+(8s)= U�G=&�1�"68G�N (& �G���+(8�

)"6-� U�=&�1����*(�(NF ��& � LG���+(8�6G1 .cM�G

R:����2 -�"H E*2& [�{/q��q/q1 EF 61 ��(,��(NF � L�"H L

E1 ����& � n���+(8���61 .

t5

� IH�56]� \�N� � �*J EK�E-6N- �&�G1?'-|�8GF& O�&�G

I6B`2){�/��s�/{2�B': � \����((& "�,- &� EFe�&[G�&

'*)2 �&"H���3NY "�(82> � �G=& W6G1�2 IH L���� ?�2 E1.

'G2 &� 56GBP2 "|�8GF& � IH ?�2 � ���$� "&[G�2 EG1 "&6G�

�&���j� � IH ?���m`= ��G�)NC !EGK��� '1H VkJ �

"6(4-��6(�� E�[k� "&[�2 � �< � '����(F�1 ?�<�EK� & �.8G- .

L�+-�GG�2 ',GG4= EP$GG% �GG��� oGGNY EGG*2& �/����/��_

'(-�= &� "�(8G2> � IH ����$G � �6- ^6$- e<�F EF 61 �(2

'2 � 6GC62 U6(��� �&�^ � IH "3 '(B�= "&[�2 E1 "&6�

& �.8- IH)�(.���8G1 ?�G<�6(F�� �6G- � IH ?�2 L�&�1�*1

3�:6� "&[�2 �1 &��> 3*(8< 'N#2 � E�:�& �-6(4-��6('"|�8GF& �

� � 56BP2�EK� � �<f'2 ��g �&�G�)����.(pH � IH�EGK�

>& ��X�1 56]� \�N� � �*J{G2 &� "H �BY EF �61 'EG1 "&6G�

u�M &�& '���BQ ?�<?'1 � ��*1�F ��G*1�F �G� � V�8GBF ?�G<

& �.8G- [G��1H Ev6J>& �3 o(,2 'F�M '���BQ �&[B� ���=

)�(EG�:�& 3�:6� � 6M EF��-6G(4-��6('eGh- "H '1�G�>�& �

�& 'GGN#2)�{.(� � �$8GG� �GGbBZ �56]GG� � �GG*J EGGK�

E*2& RB(S2 >& ?&q�/q�� "�(82> � �/�'B�2 � �G(�: � \��

� & "�(81�� EFU�=&�1 Eh./ 3*1?�&��"&��G4N< �){(LG�&

��8� "&[�2 &��> �& }���6� ��)v� EK� IH � �$��EGK�

�*J >& ��X�G1 56]G� EG�BF � _'G2 �G(�: � \�G���4�2 3G �1

)5�3C�(.IH ��6G)� "6G1 '-X6/ � IH "3 �k1�C \3Y

� ��EK� �G<)�s(G� IH '���N�G 64�[�� ���%6]GM �G1f��g

'2 �&�� .IH � 56BP2�$8� "&[�2 G1 ')�./ ?�<E'B]G� �6G/

>& "H "&[�2 � �=& ���j(2����q'G2 �G(�: � \����4�2 3G �1

)��.("68.�� �$8� "&[�2 Er-�*K E:3()2 o/�*2 � EF �F "��1

>& �GG(NF EGG1_q�GG� ��q3GG�:6� 3GG=�1 �GG(�: � \�GG���4�2

"6(4-��6(�� '2 e<�F �< 31��)�q.(EG1 E(8G1 !�G+� ?6G= >&

��)Q62 � '4�T6:6F& ?�<�"6G(4-��6( EG*�#1 �.8G- �G< �$8G� :

-�"T��GGG()N:P ( �GGG=& ���GGG$(2 ���8GGG1)�����.(nGGG:�Z

��"6(4-��6( � � �<����3GNY RGB(S2 56]G� � �G*J EGK� >&

�\6�� ���BF � �<�E8EK��& 61 =��6-� G� 'G-&�&�� � E8 8G1���

� & 6C� L���� .�.8-N:P \6��G�>& ?���8G1 � �G<�q:�!

���BFGG� E8�:�q!��6*GG�E8��:��=��6-�GG�L�GG1 E8�:s��GG�

�:���'23 �1)�_!�����.(

'BY ���$� VZ� 1 !�<E\6��� 'BF �6/ !�<3�NG= !E8G����BF !�G<

EK��� � E8��6-��= � E8��6*� �*J 3= 6GC� �& .\6��G� >& �G<

V#2 "6(4-��6(�� L��� O�P2 ?�< EGK��� ?�G< ?&EG:3()2 o/�G*2 �

3*(8< �3NY eS1 EF � &� E�:�& 3�:6� >& ?& HI2 n.= �<'3-6G

)_��q.(� �� �GGGGG*J EGGGGGK�GGGGG�\6�� ;]GGGGG� � �GGGGG<

Page 11: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

����� %&$! "��# � ����� �(�%)�*��%'� (�$� ���, -� ./�$�� ��+

{�

L)��.\]�� �+�� "��# '(���� �(�&^�T�$� _��`��0 � ��'(�%)�*��%�+�� �� �Q�, -Q� ./�$�� Q$��N%&�Q+�

.���� ��� _P%m0 ��6� � �.0'N��'C�+����� �L8�-, $� + �� .�b0 I, e$��Q0> Qp� !� ��N%&�(�Q0!> �Q

�:0 I*%]� <+� C)��� ����-� -���)P>0.05.(

0.0

0.2

0.4

0.6

0.8

1.0

1 2 3 4 5 6

��ر ���ن�� ����� ز����ن

.���� ��N%&$� ����

�'�\]��

(�&^

a

b

c

aa

b

c

c

aa

b b

ab

aa

b

c

b

a

aa

a

.���� ��N%&$� ����

0.0

0.5

1.0

1.5

2.0

2.5

3.0

1 2 3 4 5 6

��ر ���ن�� ����� ز����ن

(����\]��

T�$�

a

b

a aa

a

a

b bb

b

a

bc

a

b

c

a

cd

d

c

a

b

0.0

0.5

1.0

1.5

2.0

2.5

3.0

1 2 3 4 5 6

��ر ���ن�� ����� aز����ن

.���� ��N%&$� ����

b

c

b

a

c

a

b

d

c

ba

a

d

b

cb

a

b

a

b

c

b

c

_��`��0\]��

Page 12: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

������� ����� /������/����� ��� /�������

{�

��G� � "�(81���&�G [�.G=�*2 O'F�� l�GP: EG1 G�G-&�&�� � n'

E-6� &?�h2 � �3*( & "�(82> � ��#1 �1 E8.2 &� �BY 'EG1 "&6�

&� �� O�[� 64�N��'�1 IH n=�*2 E�.8G- l�GP: E1 a6]M

$8� - E1 ��& �.8- "T��( .6= >&?� 6GC� !�+ G�\6�� �G<?

$2�& � 3�L��EK� %6]M �1 �:X �F n=�*2 ���$'� IH

"�(81�� ;]��&.

� ��EGK� �G*J 3GG= GG�\6�� i*GGC �>& �G<Cyclotella �

Surirella �*�(GG=& �GG1 56]GG� \�GGN� �?VGG#2 "�(8GG2> �GG��� L

"&�&�� ���3-61 L.2 &� �BY'&6� %6]GM EG1 "�F ���G$'!IH

1E�2 a6]M?&�G1 "�(8G2> � EGF & �.8- IH ?� 3G �

�4��81 ����?� >& �"6(4-��6( - n=�*2 �<�-&�&�� � �8'e<�F

EbJ�2 ;1�Q &?2'�3G1�)�q!����{.(6G= >&?�e<�GF !�G+

2�-&�&�� "&['�3J �� �?F�� �EG-6� n&?G�\6�� ��G#1 � �G<

2'�]2 ;%�J 3-&6� "6G(4-��6�> OG=6� "3 &�K � }� �G<

(J'<�2 ��X �� "��3 �1 �&6M "6(4-��6()��.(

��"6(4-��6( �<?���BF ���� � "�(81�� !��#1 � E8�F�� [G�n

E-6� &?�h� ���. ��82 ?� & .G2 &� �BY'F EG1 "&6G��$G�IH �

��&� � EK����G1�1 O?���BF G�& �.8G- E8)�q!�����.(

1EBF�6/'81���?>&A.BC �<?���BF �&�& E8?�:6B= ��&6'

[:6B=?81 �3J �� EF 3*(8< ���?B1�Q �"3 }�]2 � &�K �

�h2 � &� �#-H��= �1 E8�E-6� ��<? 3Q�� �[:6B= ��&6?.Q >& G�;

�E26�� 2 �3P2 �<'�N-�3)�q.(6= >&?�&[�& �+�-&�&�� e'

���BF�2 &� ��#1 � E8'- }�]2 E1 "&6� �4� � "3, G� :6� � �G�3

E-6� ;�2 �<?& >& �A.BC L81 EF �<�� �� G� 3*(8G< [)EGBNC >&

Chlorella ( & �.8- .N= E(= >&�V#2 !�<3 ���Li*GC &G�L

��EK� Cosmarium IH "3G �G= �G1 EGF �=& )�G2?�/�

(-�= EC�'"�(8G2> � &�� (� ��EGK� �3G� 3,G- )�q.(>&

������6*�E8E-6� �G<'�i*GC >& Ceratium 56G/ \�GN� �

� � 5�=�/ � �#-H e*F&�� EK��& �& 6C� EK�';]� �<

& E1 �3P2�%�M ��+(8'>&��EK� �� �3.2 "�N� �2'*F�EF V

*K >��1 ��&� L�`'!�2 "&[�2 ^6$- �bBZ !�6- &62 !?�Gj2

V@&�G� "��3G**@&�K )"6G(9-��6�> �G<(!OG1&�� '(8G�-6��(-H �G1

"�<��� !?[1H ���j� � IH �&6G<!?T6:��3G�< � IHoGNY IH

2�3 �1 �g)�{(.

1��(,�p6*� L>G�(8'��"6G(4-��6( �G< & � G�hP� LG�� o

& � "�(81�� � ��#1 56]����+(8�61 .X EBNC >&�;:�N(J& '

&[�&�2 p6*� e'�G2 "6G1 n=�*2 E1 "&6�?��X ��$(G=& !IH

<�2GG�4-��6�> >& "�"6GG( e<�GGF � �GG< .8GG-'&�GGK ��,GG� ?

-6GGG(4-��6�>'!�&62 ��� -3GGG)2'��GGGj2?>&���?IH

��� EK�;P2 &G���+(8 �G<?�& �.8G- )�!�!_!�q!��!

�� !�����.(& � GG���+(8 �oGGNY ��EGGK�u3GG-& !IH� \&�H

� V/�� "�31&�&?, 6� '>&��[1H "�<�?E/6Z ���*G � ��

2 IH �'3 �1 .�IH�<?��EK� &?�1>& �G(NF oNY �q�G(2

e<�GGF "3GG W6GGBS2 oGGNY GG-&�&�� e<�GGF nGGC62 IH'

��-6(4-��6('2'�� .6= >&?�N<& !�+ G� "6G(4-��6�> ��G<

F�� 5�GG(*F �GG�n!GG-&�&�� '�>?� �6GG� �-6GG(4-��6('�

��EK� �<?1H 01�*2 �'��2�F G2 cS,G2 '3G �1 >G�&�&�GK ?

"6GG(4-��6�> � >& �GG<�"6GG(4-��6( > e<�GGF �GGY�1 �GG<?�6GG�

��"6(4-��6( 2 �<'(J 6 '- � �$8� �bBZ ��& �56BP2 "T��(

3 �1 X�1 V<)�����.(&�1�*1�!L� p6*��"6(4-��6( � �< � ��G#1

� "�(81���h2���� �1 E8�"�(82> � [1G�6G1 �(, .6G= >&?��G+

2 "�N�'EF �� E=�2 n=�*2 �(81 6C� &?* �'�3GJ �� ?

&[�& nC62�= e�BG�8�� IH V�G= ��F�� �$2 ��G. G� �(8G1 >& 3

2'& � EF 6 ���+(8 �<?�G(NF "�G42& ��X�1 oNY �1 ?6GC�

�&)�q.(�h2�& E8���+(8 p6G*� l�P: E1 �< ��G� 56]G� ��� [

"�(82> EF & "�,- 1��(,�& � p6G*� LG���+(8s����EGK�

�=& .V#2 ���: L�;& � p6*� "61 VF �56]� L�h2 � ��G1 E8

56]���+2 &� '�3P2 E1 "&6���G2 �'�IH& � G�56]G� L

-6GGBF � 3GG � nGG=�*2 EGGF & �.8GG-'8GG1 � "3GG ���?>&

E-6� �<?��-6(4-��6('-��8)�q!����{.(�;]� "�(8G2>

���6@ "3 56/ >�� ����j� E��&> �6- 3� �6M � x�GY ?�G<

��G��&�jC'OG=6(2 �X�G1 'G2 3G-&6� ;G2&6Y>& �3G-�&>�1 3G �

D.BC �<�3�:6� c:�M �#-H !3 �1 1E?�6/ E@O�&� ?&�1 3G �

i*C ?�< nG:�Z 3Y�8G2�- �3G �V@&�G� �G#-H VG@ 'G2 6G �

Page 13: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

����� %&$! "��# � ����� �(�%)�*��%'� (�$� ���, -� ./�$�� ��+

{{

i*C ?�< I6Bj2 V< '2 3*-&6� �� �>&3-& ?&6M &� "�,G- 3G*<.

E1 'B@�6/ EN< ?�<�6(@�� '���N� 69�[�� �(8�>'�G1 nG�@��

E-6� ?&�>?�6� "6(9-��6(�� �<��gf� E( &�� ��&3h2 �p6G*�

�#-H &� ���j� '2 3<.

��ZN��^��

>&���NJ O�GP2 �G��$J ;GF ��&& ':�G2 ?�< "�(G=& �8G�>

��/ n:�Q � "�#$%&��s�{�!G*� � ',G<�|� �-��)2�?��

� "�#$GG%& '()*GG% ��+,GG-& 'GG)�./ 01�GG*2 �39,G-&"X678GG2

� "& �&�Q ���(M& � ?&�1 "�(=& IH "�2>�=f�G*J 3G= ��8�=

��F ?&�C& L�J �'2 '-&�3Q � �9,� V��N-.

G��0 ���0 ���1%��

�.3GGNJ&?\ !.�.��66.GG=��1'��"6GG(4-��6( �GG<?��IH � "62�GG< EGGK� �GG<?GG �J �"H E../ 01�GG*2 EGGBk2 �GG)'&GG�"&�

�� :���6.

�.� !&�H A.= .�\.��42'2.���� ."6(4-�� '-&�&�� � e*F&�� 5�= '/ ':[-& IX�� � �#-H eh- � �< ?�G<���6�G� ����.EGBk2

"&��& ��� 'NBY��)�:(������.

�.2�Z'p !.!�.�#(C& ?�}.�&> V=�Q .��_s .=��1'E-6� p6*� &?T6:6F& ��� A�"6(4-��6( �<?��"�G+-[1 EK� .GNBY EGBk2'

�& ���"&��:���s�.

s.J��'\ !.��{_.-6(4-�� ��&[� '< �T��� �T6:��3?< �G�1��3�T6:6?G:[-& IX�G� '.hP� [GF�2 �G ��Gh���'G� "�(G=& �!"�

���a.

�." !'-���6% '16.P2 .��__.\61 ��):�`2 ��&[� � �1 �*J p6*N2 ��4 '1X�� � '-�(8<6F o/�*2 '(M�* f�h��& "�42& 3�F�EGh`*2

g � �3 ���$J�82&� "6�8-&6*F � �. .��K �= �.

�.-�C3NP2'W !.�p.J��3?.����.�=�* '���"6(4-��6( �<?���?�#-H e*F&�� � �[M.hP� "�2>�= �G �>6G2H � ��Gh���

&�hP� [F�2 !"&��� ��h�!"���a.

7. Auer, M. T., M. S. Kieser and R. P. Canale.1986. Identification of criteria nutrient levels through field verification of models for phosphorus and phytoplankton growth. Canadian Journal of Fisheries and Aquatic Science 43: 379-388.

8. Barsanti, L. and P. Gualtieri. 2006. Algae: Anatomy, Biochemistry and Biotechnology. CRC Press, Taylor and Francis Group, 320 pp.

9. Batelle Columbus Laboratories. 1971. Water quality criteria data books. Volume 3. Effects of Chemicals on Aquatic Life – Selected. Data from the Literature Through 1968, for the U.S. Environmental Protection Agency, Project No. 18050, Battelle, Columbus, OH, 530 pp.

10. Bellinger, E.G. and D. C. Sigee. 2010. Freshwater Algae: Identification and Use as Bioindicators. John Wiley & Sons Ltd., 271 pp.

11. Benndorf, J., D. Uhlmann and K. Putz. 1981. Strategies for water quality management in reservoirs in the German Democratic Republic. WHO Water Quality Bulletin 6: 68-73.

12. Boyd, C. E. 1979. Water Quality in Warmwater Fish Ponds. Auburn University, Craftmaster Printers, Opelika, Alabama, 359 pp.

13. Chaghtai, F. and S. M. Salfullai. 1988. An Illustrated Account of Species Ceratium, University of Karachi, Pakistan, 50 pp.

14. Clesceri, I., A. E. Green berg and M. A. Franson. 1998. Standard Methods for the Examination Water and Wastewater, American Public Health Association, Maryland, USA, 1368 pp.

Page 14: ˆ˙ ˝ ˜ ˛ ’˚2 450 3 6˛ ˜78-2 $ ˜ -’, · PDF file˜8g1 ˛3g2 >& ˛& g˜˜j lp: e1’‘˜p2 8> ˛˚4,2 ... n gg . gg ˝ \ggk-& ( ˙ ! q) ’gg=* ?

������� ����� /������/����� ��� /�������

{_

15. Cox, J. E. 1996. Identification of Freshwater of Diatom from Live Material. Chapman and Hall, London. 16. Davis, C. C. 1955. The marine and Fresh-Water Plankton. Michigan State University Press, 562 pp. 17. Day, J. W., C. A. S. Hall, W. M. Kemp and A. Yanez-Arancibia. 1989. Estuarine Ecology. A. Wiley-Interscience

Publication, John Wiley and Sons Ltd., New York, 558 pp. 18. Demirbas, A. and M. F. Demirbas. 2010. Algae Energy: Algae as a New Source of Biodiesel. Springer Pub., 199 pp. 19. Demirbas, A. 2006. Biogas potential of manure and straw mixtures. Energy Source A 28:71–78. 20. Gibson, C. E. 1997. The dynamics of phosphorus in freshwater and marine environments. PP. 119-135, In: Tunney,

H., O. T. Carton, P. C. Brookes, A. E. Johnston (Ed.), Phosphorus Loss from Soil to Water. Cab Intl., Harpenden UK., 467 pp.

21. Krebs, C. J. 2001. Programs for Ecological Methodology. 2nd ed., Benjamin Cummings, 624 pp. 22. Lee, R. E.(ed.). 1989. Phycology. 2nd ed., Cambridge University Press, 645 pp. 23. Margalef, R. 1958. Temporal succession and spatial heterogeneity in phytoplankton. In: Buzzati-Traverso, A. (Ed.),

Perspectives in Marine Biology. Berkeley, CA, University of California Press. 24. Marsden, M. W. 1989. Lake restoration by reducing external phosphorus loading: the influence of sediment

phosphorus release. Freshwater Biology 21: 139-162. 25. Odum, E. P. 1971. Fundamentals of Ecology. 3rd ed., W. P. Saunders Co., Philadelphia, London, Toronto, 574 pp. 26. Omori, M. and T. Ikeda. 1984. Methods in Zooplankton Ecology. John Wiley & Sons, New York, 332 pp. 27. Onyema, I. C. 2008. Checklist of phytoplankton species of the Iyagbe Lagoon, Lagos. J. Fisheries and Aquatic Sci.

3: 167-175. 28. Prairie, Y. T., C. M. Duarte and J. Kalff. 1989. Unifying nutrient chlorophyll relationship in lakes. Canadian

Journal of Fisheries and Aquatic Science 46:1176-1182. 29. Redfield, A. C. 1934. On the proportions of organic derivatives in sea water and their relationship to the

composition of plankton. PP. 176-192. In: James Johnston Memorial Volume, Liverpool University Press, Liverpool.

30. SPSS, 2002. Statistical Package of Social Science. Ver, 11.5. SPSS, Chicago, IL, USA. 31. Uhlmann, D. 1958. Biological self-purification in sewage ponds. Verhandlungen des Internationalen Verein

Limnologie 13: 617-623. 32. Uhlmann, D. and E. Albrecht. 1968. Biogeochemische Faktoren der Eutrophier-ung von Trinkwassen-Talsperren.

Limnologica (Berlin) 6: 225-245. 33. Zar, J. H. 1984. Bioststistical Analysis. 2nd ed., Prentice Hall Inc., Englewood Cliffs, New York, USA, 718 pp.