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K
Koninkrijk der Netherlanden
WWF Mongolia
2009
UND
DP M
M
Pub
lic D
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e A
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rized
Pub
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Pub
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Pub
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Pub
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Pub
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Pub
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Pub
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: : ,
- 2010 1-
25-
:
:
:
:
(PhD) .
Rich Harris (PhD)
C. ( )
.
.
. (PhD) .
. . . . . . . .
2009
,
,
: - 2010 1- 15-
.
: 2010 ... …- .
2009
2
2010
(Ovis ammon L.,1758), (Capra sibirica L.,1758)
,
: . (Ph.D)
: .
: Richard Harris
2010
2009
3
............................................................................................................... 4
................................................................................................ 5-8
........................................................................ ............................................ 9
, ........................................................................................................ 10
.............................................................10-20
..................................................... 20-21
....................................................................... ……………… 21-22
……………… 22-28
, ( )............. 28-109
- ..................................................................................... 29-45
..................................................................................................... 45-58
.................................................................................................. 59-63
- ........................................................................................ 63-75
........................................................................................ 75-79
.................................................................. ……………...79-81
......................................................................................... 81-91
.................................................................................... 92-93
..................................................................... ………….. 93-98
..................................................................................... 98-106
............................................................................................. 106-107
.................................................................................. 107-108
.................................................................................. 108-109
.................................................................................. 110-113
.......................................................................................... 114
.................................................................................. 114-115
........................................................................................ 116
.................................................................................. 117-121
.......................................................................................... 122
2009
4
, , -
, ,
- , ,
“ ” , “ ” ,
- , , - , , , , ,
, , , , -
.
-
. , . ,
Dr. Matt Reeves .
,
- .
2009
5
,
(Ovis ammon), (Capra sibirica) , , -
, , - 2009 10- .
2009 3, 10- 2 11
12 21 . , , ,
, , . 2001
,
. , .
1:40,000 1:50,000 .
. GPS ,
, . 1 3
100 15 . GIS-
. distance sampling .
. Distance ,
. .
50215.4 2 46,603 2 158 10-20 20 , 880 ( )-
.
. ,
. 50215.4 2 95%-
9193-43135, 18140 ( -1).
1- . , 2009 10 .
,
95%
95% 572 444 927
- 2123 931 3761 / 2913 1361 4967
2009
6
2338 1505 15408
- 1556 1066 9158 2311 341 3400
2404 1198 4852 1756 1160 2368
834 417 1664 1033 591 1591
40 23 50 50
180 30
18140 9193 43135
162 2541 (3- ). ( ),
95%- 36018 (13840-43873) ( -2).
2- . , 2009 10-
A
(2009, 2)
1 127.86 100 2 - 7522.3 5.15 3874 3 5649.6 5.15 2909 4 - 8917.6 5.51 4913 5 225.5 5.32 120 6 1564.7 9.70 1518 7 6078.4 2.20 1337 8 2568.9 1.30 334 9 5309.5 25.09 13324
10 80.9 1.85 15 11 1335 14.30 1909 12 2.1 0.00 42 13 3709.8 2.94 1091 14 13021.5 3.48 4532
55985.8 6.45 36018 (13840-43873)
. ,
(Method of Distance)- .
95% .
2009
7
, ,
, . 2001
, , , ,
.
, , . ,
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. ,
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. 2009 3
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. ,
, .
2009
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. , . 2009 3- -
, ,
2
, .
, , ,
, ,
, .
, ,
Distance Sampling, - ,
.
2009 9 16-17-
, , ,
,
, GPS- , ,
, 1:40,000, 1:50,000
.
, , ,
.
2009
11
.
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. 2009
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.
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. ArcView
. ,
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,
.
,
.
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.
. - , , , - 1:40000,
1:100000, -
1:50000 .
, , ,
. 2000-2002
Enhanced Thematic Mapper (ETM)- . 200 -
2009
12
Shuttle Radar Topography Mission-
.
. ,
< 1.0 .
. :
1) Distance sampling
,
2) -
Huggins closed-capture (White, 2008)
( ),
.
, ,
GPS, .
100 3-4
1 .
.
.
. ArcGIS (ESRI,
Inc.) ArcGIS V. 9.2-
.
. 3-5 . 3 -
Distance
Sampling . 5
.
.
,
.
2009
13
.
, > 85%-
, 95%- . Distance Sampling
,
,
.
,
.
. - 2009 10 12-28- 2-4 2
11 9- 1:40,000
. ,
Distance sampling (v6.0), Release 2
(Thomas et al. 2009) .
.
< 3 - . ,
Multiple covariate distance . 2- . -
( x)
.
2009
14
. 3 1 . 2247 2 10
1532 2 5 1:40000
. , -
2 .
3- . ( x).
. 2-4 2 2009 10- 5-22-
15 – 1:40000
. 7
. 3
.
. - 2-4 2 15 12-
1:40000 .
,
distance sampling DISTANCE 6.0, release 2 (Thomas et al. 2009)
.
. < 3 -
. , Multiple covariate
distance . 3 .
2009
15
4- . - ( ).
. 1-3 3 , - 3
, - 3 . ,
. (ScD) .
.
. 3 1 11 2
.
Distance Sampling .
, . 5- . ( x).
2009
16
. ( a ) 2-3 3 1:100,000
12 8- .
, 50-60, 10-20 ,
.
– , - ,
.
Huggins closed-capture
. ,
.
6- .
( x).
. 2009 10 9-28- 2-4 2
31 23- 1:50,000
. ,
distance sampling DISTANCE 6.0, release 2
(Thomas et al. 2009) .
.
< 3 - . ,
Multiple covariate distance .
2009
17
7- . ( x).
. 3 1 24 21-
. 3
. , ,
, Laser Range Finder
distance sampling . DISTANCE 6.0, release 2 (Thomas et al.
2009) .
. < 3 -
. , Multiple covariate
distance .
8- . ,
( x).
2009
18
. 3-4 1 21 18-
. 3 . , , ,
Laser Range Finder distance
sampling . DISTANCE 6.0, release 2 (Thomas et al. 2009)
.
. < 3 -
. , Multiple covariate
distance .
9- . ( x).
. ( )
, 2
. , 2000
, ,
.
3 .
,
.
2009
19
K. . . 1 , ,
20 12- .
.
K. . . 3 1 . ,
, , , , , ,
. ,
. (Index)
.
K. .
~ 10 12 . 24
- .
.
Geist (1991), Shackleton (1997), ,
IUCN (2008)
(O.a.darwini), (O.a.ammon) 2
. ,
.
2001
. 1980-
-
. 1970- 50,000,
1985 ( 1970, 1994, . 2002b, Zahler et
al. 2004, Clark et al. 2006, ) 60000
. (1997, 1994) 20,000-
. 2001 -
,
47,898 km2 13,000 – 15,000 2009
, . 2001
10-20 ,
, ,
.
2009
20
. 2001 1976
- , , – , ,
, , 7 34
31,2% .
, -
62,5%, 53,3%
( , 2001).
( , 1987)
1986 80,000 1980-
(Fedosenko and Blank 2001) . Mallon
(1997) , ,
.
.
(Ovis ammon),
(Capra sibirica) . 2
, .
“ ”
. 1953 . ,
(CITES) II , 80
. (O.ammon)
1987, 1997 “ ” “ ”
(Clark et al. 2006). 2
. 2,6
.
, ,
. ,
.
, ,
2009
21
.
,
. ,
.
,
, ,
(Zahler et al.
2004).
23.2%
9.36% ( , 2001).
14% , , ,
, , , ,
, , , -
( , 2002b).
2009
22
,
50215.4 2 .
10- 3%-
.
. , , ,
- .
.
2001 2009
. ,
9,7-8,9 .
Home range .
.
, -
1:500000- , ArcView
.
.
3,2 , 2,3
, , , - , 1,2-1,9
.
, , , ,
1,1-2,3 .
, -
(3- , 10- , 1- ).
2009
23
3- . ,
( 2)
(2001, 2)
(2001)
(2009, 2)
(2009)
-
( 2)
1. - 35663.08 2274.82 6.38 4244.3 11.90 1969.52. 62060.33 2955.35 4.76 3560.8 5.74 605.53. - 84714.59 6263.84 7.39 6465.1 7.63 201.34. 5498.32 9.19 0.17 29.0 0.53 19.85. 97314.96 8315.33 8.54 9671.9 9.94 1356.66. 56783.69 3991.22 7.03 1723.5 3.04 -2267.77. 22949.22 1184.74 5.16 1184.7 5.16 0.08. 34925.78 3716.55 10.64 2151.2 6.16 -1565.49. 164134.80 8990.77 5.48 6371.7 3.88 -2619.110. 16456.82 120.01 0.73 1070.9 6.51 950.911. 53173.00 1296.91 2.44 2992.0 5.63 1695.112. 16597.61 694.12 4.18 1315.9 7.93 621.813. 19298.53 63.04 0.33 613.8 3.18 550.814. 65889.57 8028.58 12.18 6369.0 9.67 -1659.615. 16625.57 828.12 4.98 720.6 4.33 -107.516. 18448.90 0.00 0.00 1327.1 7.19 1327.117. 4974.20 0.00 0.00 403.8 8.12 403.8
752085.85 48732.59 6.48 50215.4 6.68 1482.8
2009
24
10- .
1- . (2001-2009)
95%-
9193-
43135, 18140 (3- ).
10- .
2001 2009
.
.
2009
25
. ,
, .
.
2009 - , -
226,6%-309,5%- , , , 122,4%-175,1%-
. , , 58,9%-
97,2%- .
, , ,
. , 40,5%-
63,6%- . ,
(4- ).
4- .
(2001)
(2001)
(10 2, 2001)
(2009)
(2009)
(10 2, 2009)
1. - 2274.82 650 2.9 4244.3 2123 5.02. 2955.35 290 1.0 3560.8 572 1.63. - 6263.84 380 0.6 6465.1 1556 2.44. 9.19 85 92.5 29.0 140 48.35. 8315.33 1745 2.1 9671.9 2773 2.96. 3991.22 850 2.1 1723.5 2338 13.67. 1184.74 110 0.9 1184.7 40 0.38. 3716.55 1840 5.0 2151.2 1756 8.29. 8990.77 4040 4.5 6371.7 2404 3.810. 120.01 5 0.4 1070.9 50 0.511. 1296.91 375 2.9 2992.0 834 2.812. 694.12 380 5.5 1315.9 1033 7.913. 63.04 30 4.8 613.8 30 0.514. 8028.58 2460 3.1 6369.0 2311 3.615. 828.12 175 2.1 720.6 180 2.5 13 86
55985.9 2 .
1:500000-
1:100000-
2009
26
.
, , , -
.
- , - , , ,
. 83,1%
.
2 .
. “ ” (1970,
1980)
. 5- , 2-
, 11- .
5- .
A
(2009, 2) 1 - 10 7522.32 12 5649.63 - 8 8917.64 4 225.55 5 1564.76 11 6078.47 3 2568.98 7 5309.59 1 80.9
10 7 1335.011 1 2.112 4 3709.813 13 13021.5
86 55985.9
2009
27
2- .
11- .
162 2541 .
( ),
95%-
35373 (13840-43873) (6- ).
2009
28
6- .
A
(2009, 2)
1 127.86 100 2 - 7522.3 5.15 3874 3 5649.6 5.15 2909 4 - 8917.6 5.51 4913 5 225.5 5.32 120 6 1564.7 9.70 1518 7 6078.4 2.20 1337 8 2568.9 1.30 334 9 5309.5 25.09 13324
10 80.9 1.85 15 11 1335 14.30 1909 12 2.1 0.00 42 13 3709.8 2.94 1091 14 13021.5 3.48 4532 55985.8 6.45 36018 (13840-43873)
3- .
2009
29
. -
A. 1.
92.3% 4,749 2 60
. 59 .
41 500 . 9 7-
. 2-76- ( x = 12.5, SD = 13.8).
424 , 291 , 30 , 93 , 10
. (> 2 )
,
.
3 (hazard rate key function with Hermite
polynomial adjustments, hazard rate key function with a maximum of 2 cosine
adjustments, half-normal key function with maximum of 2 cosine adjustments)
2,627 (SE =
212 ).
- . . . 2400-3700 ,
2600-2900 , 2700-3000 ,
- 2700 – 3300 , 2600-3800 , 2200-
2600 . - ( , ), ( ,
, , , , ,
, , ,
, , , , , , ,
, , , , ,
, , ), ( , ), ( , , , , ), ,
( , , , , , ,
), ( , , ,
, , , ,
, , , , ,
), ( , , ,
, , , , ,
, , , ,
2009
30
), ( , , , ,
, )
( . , . , 2004).
1991 - 6000 .
, 2001 -
2408,14 2
( , 2002).
- 2008
10 6333.4 2 ( ,
2008).
12
4825.55 2 .
3511.5 2 , 466.23 2
.
847.82 2 ,
/ 2/.
4- . -
2001 1991
59.9%- /4- /.
- ,
2009
31
, ,
.
2009 2001
39.5%- “ -
”-
, .
, ,
.
- :
“ ” . -
- “ ” ,
, , , ,
,
466.23 2 .
, , ,
,
( . , . , 2004). ,
“ ” , ,
.
“ ” .
“ ” ,
, , , , , ,
, , , , , ,
, , , , ,
, ,
. 598 2
.
.
, , , , , ,
, 538 2 .
2009
32
8 , ,
, 4 16
100 . 10- 7-
- -
.
(125;121;100 )
“ ”, “ ” ,
.
.
, , , , , , ,
,
376.3 2 .
1992, 1993 ,
( , 2007). 2007
“ ”
,
.
. 296 2
.
. -
- .
,
. ,
,
.
. -
, , , , ,
426,4 2 .
2009
33
,
,
.
. , ,
299.62 2 . ,
30
2-3 .
. ,
-
264.81 2 .
2 .
(1989 )
( , 2007).
.
90.12 2 . 6
. .
,
.
, , ,
. ,
.
12- . -
2009
34
, “ ” ,
, , , 6
41 505 . 425
(84.1%) .
5- . -
: : : 4.8:19.9:6.9:1 .
34.7% .
60
2009
35
. 2008
2
7 . 2008-2009
, ,
.
- , - , - 1:4.1, 2.8:1,
19.9:1 2001
.
- , - , - , -
- 1: 1,5; 1: 0.3; 1: 0.2 .
- 2001
4 .
: .
2001 .
( , ),
.
, ,
,
.
6- . -
-
3 2007
. - ,
,
2009
36
,
.
2.1%:65.9%:34%
. 2007 12-
, ,
.
.
( . , 2004)
.
, 2007
, ,
, ,
/6- /.
,
.
,
. . 2007 12-
. -
. ,
.
.
-
51.5%, 51.7%, 16.7%-
.
,
, ...
, , - ... , ...
. . (1990) .
1991 , ,
, /1 . - 0.01-0.05 /, , ,
/1 . - 0.05-0.1 /, ,
, , , , , ,
2009
37
, , , / 1
. - 0.1-1.0 / . -
6000 . 2500
/ , 1992/.
2001 -
2408,14 2 710
( , 2001).
- 2008
1000 - 0.32 2382
( , 2008)
- ,
,
.
7- . -
, 1999 2003 2007
1000 -
1000 -
1000 -
1
300-450 7.8 650 17 - 741±8 0.52
2
- - - - 125±12 1.16
3 1800 14
- - 227±16 1.6
4 - - - - 5 - - - - 100±15 0.53
,
.
,
,
. : 2007 12
100,25±15, 4
( . , . ,
. , 2007).
. (2007) 2
. .
2009
38
- 2009
0.447 / 2 (95% 0.196 – 0.792), 4,749 2
2123 (95% 931 - 3,761) .
7- . -
- 2009 1991
15% .
, ,
2001 66%- . -
- ,
,
.
,
.
, , , ,
, , , ,
. , ,
, , ,
.
- ,
2009
39
.
.
- 2008
15.4%- 7097.7 2 ( . ,
2008).
- :
- ,
, , , , ,
;
- , ,
;
- , , ,
, , , , ;
- , , , ,
, , ;
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, , ;
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, ;
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, , ,
;
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, ;
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“ ” , , , ,
, ; , ,
.
,
.
2009
40
, ,
.
, ,
.
13- .
,
,
,
, . 13
249 . 114 (45.7%)
.
2009
41
8- . -
: : :
3:38.5:1:14.5 .
2008 21.8%- 2009
1.8
, 90
/8- /. -
2008 100 62 2009
2 100 37 .
50 . 2008
, ,
, , ,
,
. 2
2009 ,
.
2009
.
, ,
,
. - ,
- . /1976 / ,
2009
42
/1975/
( , 2002).
- 1991 - 5200
( , 1992).
2008
- , , , , ,
, 7097.7 2 3786
( , 2008).
13 249
6 ,
.
- 3874 .
2
.
- .
,
,
. .
.
.
.
.
. 2
. ,
. ,
, . -
, .
2009
43
-
,
, , , , ,
, ,
, - ,
, . , 2001 -
1991
59.9%- , 71.6%- . 2001
, ,
,
.
, ,
,
.
.
, , ,
. -
2008 ,
.
, .
2008, 2009
16.9%- , 35.2%-
, 100 18.5%- , 86%-
. 48.3%-
92.3%- .
, .
2009
44
2008 ,
.
, ,
. , “ ”, “ ”
.
, 2004 -
30 , 12.0-16.0
, 2000-3000 ,
12 , 5.8-7.5 , 1300
. - , ,
, 3- , 11- .
(
, 2007).
“ ” ,
, , , ,
1,2,3-
.
. , , ,
, . , ,
,
( . , . , 2007., . , 2008).
“ ” , ,
, , , ,
,
(600 , 15.0-18.0 )
- .
,
“ ”
( . , 2008).
, ,
2009
45
,
. ,
.
,
. ,
, ,
,
.
,
. 2007
.
1989
, 2007, 2009
.
,
.
, , , ,
. 1989 2006 60
56 , 56 38 .
56,2±4,66 / , 2007 /.
.
. . 1.
19 414 . 5 3-
. 1-76- ( x =19.4, SD =
21.9). 164 , 89 , 15 , 11 ,
49 . - 391
(2009 7 23-28) 315- , 134 , 95
, 19 , 67
217 .
2009
46
- 1991
,
, , ,
,
. 2001
6 , ,
, ,
,
( 2001).
- ,
, , ,
540.82 2 .
, , ,
, .
, ,
.
-
, 86.56 2
.
- 146.61 2
. , , , ,
, , .
. ,
, .
9-11- , .
- . 12.22 2.
- 235.27 2 .
, , , , ,
, , .
,
2009
47
. , , ,
.
- 74.87 2
- . 15.85 2
- , . 56.86 2.
- . 33.16 2
- , . 69.47 2.
- 7 880 .
, 2001 706 2 ( , 2002).
, ,
,
, , ,
, , ,
, ,
, .
2009
10 1271.69 2 .
1022.14 2 ,
, , ,
249.55 2 , .
9- .
2009
48
2001 1991
19.8%- /9- /.
,
, ,
.
2009 2001
31%- , 2004 ( , 2004) 25%-
. ,
, “ ”
,
.
, ,
.
14- .
, , , ,
5 18 414 .
164 (39.6%) .
10- .
2009
49
: : : 1:11.6:1.4:6.1 .
11.8%
. 50
40 .
- 8:1 /10-
/ .
: 2004
2.7:1 16.1%- 2009 1.9:1
30.4%-
, .
- , - , - 1:11.6, 8.4:1,
1.8:1 2001
. - ,
- , - , - 1:1.4; 4.1:1; 2.1:1
.
,
.
- 2001
11 .
2001 51.2%- ,
14.3%- .
2009
50
. “ ”
.
, ,
, ...
, ... . . \ 1990 \ .
- 1991 260 2
0.1 , 188 2 0.3 , ,
, , , 64-112 2 0.05
450-500
.
2001 706 2 1000 - 5.3 380
( , 2001).
2004
767 2 681-691
( , 2004).
,
,
.
” ”
,
/11-
/.
11- . “ ” -
2009
51
, , , ,
5 18 414 .
5 3- . 1-76-
( x =19.4, SD = 21.9).
12- .
- 2009 7
2247 2 591
Rich Harris .
2001
,
.
2004
,
. 2009 2004
13%-
,
.
2009
52
.
591
.
.
.
1315,9 ² .
24 7 93
414 591
.
multiply survey 4 -
25,14
² (S=2 R) .
527,52 ² 414 .
1315,9 ² [=1315,9*414/
(2*3,14*4*21)] 1000±33 .
,
.
.
:
- 89.70 2;
- 150.16 2;
- 50.89 2;
- - 68.33 2;
- 219.91 2;
- ,
122.18 2;
- 61.83 2,
98.56 2, 120.20 2, 28.19 2,
47.87 2;
2009
53
- 254.32 2
.
1335.0 2
,
.
15- .
,
1 78
.
13- .
2009
54
2004
: : : 1.2:3.8:1:1.7 .
/13- /.
- 2004 100:45
(90%) .
- 26% /
, 2004/.
, ,
2004 , -
, .
3430 2 8972
( , 2004).
, 2009
, - ,
.
8- .
/ 2/
/ /
1
150.16 800-1000
2
89.70 50
3
68.33 200
4 ,
122.18 300
5
219.91 2000
6
50.89 100
7 , , ,
,
356.65 620
8
254.32 500
1312.14 4570-4770
76 1 .
, , ,
2009
55
, , , , , ,
o
1:500,000 .
1335 2 1900
.
, ,
( , 2008). ,
6 , 7
. , 2001
1991
19.8%- , 15.6-24%- . 2001
, ,
,
. 2004
2001 8%-
45%- 2 .
, .
2004, 2009
79.8%- 100 28.1%-
20.7%- , 32%- .
.
, .
, ,
.
2009
56
,
,
.
,
.
, , , ,
. ,
.
, ,
.
, ,
,
.
,
, ,
, ,
.
,
, .
2007 217255
, 2005 199733 -
. 2008 2004
893 - ,
1472 - ,
. 2007
2003 24 ,
24 , 92 ,1 ( , 2004) ,
.
, .
. (6.7%)
2009
57
.
,
, ,
.
, ,
,
.
,
,
,
,
.
1. , ,
,
.
“ ”
, .
,
,
. “ ” - 2009
2003 72.5%- , 2007
51.2%- ,
.
,
2009
58
, .
.
2.
.
, , , GPS, , ,
.
.
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4.
,
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,
6. , , ,
7.
,
,
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.
. , ,
.
.
.
. 2
.
2009
59
. .
80%-
, 14 272 .
216
341 . 5 ,
2 .
33 780 .
1246
2200 . 4 ,
2 .
.
1992
.
,
, , .
16- . , 2009
2009
60
17- . , 2009
9- .
1 1992 3000 5000
2 2001 2000 - ,
3 2007 476 4191 , , “ ”
4 2008 690 3444 “
”
5 2009
10
2311 4532 , “
”
, , ,
.
.
341 .
2009
61
2001
.
Richard Harris- .
3172,6 ² .
2 10 11 183 .
multiply survey 4 -
18,84 ²
(S=2 R) .
251,2 ² 272 .
3172,6 ² [=3172,6*272/(2*3,14*4*10)] 2300±11
.
, ,
4 144 39
27.1% , 60 41.6% , 23 16% ,
22 15.3% .
: : : 27.1%:41.6%:16%:15.3% .
1:0.6 : 1:0.3,
: 1:0.4 .
, , , ,
27 1273
.
. 5-128 (10- ).
10- . (2009 10 )
1-10 10-20 20-30 30-40 40-50 50-
3 4 1 1 1 3
9 51 24 36 49 259
2.1% 11.9% 5.6% 8.4% 11.4% 60.5%
, ,
2009
62
,
.
, , ,
. , , , ,
, .
, ,
.
, ,
,
. ,
, 2000-2003 4
, 2001-2002 .
, .
:
o .
, 2002-2004
. .
o
, .
o ,
, , .
1990-
.
,
.
.
,
2009
63
.
1992 ,
,
.
, . , ,
, ,
, , , 3
163 , 2 115 , 2, 2
.
,
.
,
, ,
. .
. -
12162 215 12 72
( 3-10) 85.8% . 50
.
16 83 . 12 6-
. 1-13- ( x =5.2, SD = 3.4).
83 , 38 , 20 , 16 , 9
. (> 2 ) ,
.
2 (hazard rate key function with a maximum
of 2 cosine adjustments, half-normal key function with maximum of 2 cosine
adjustments) ,
. 1,236 (SE = 282m)
0.321 / 2 (95%
2009
64
0.137 – 0.753), 12,162 2 1556 (95%- 1,666 - 9,158)
.
2001 -
6 6263,84 2
,
69.5% .
-
. , . 2007 -
27.3%-
.
2009 ( “ ” ),
, ,
, ( ) ( “ ”
) , .
,
.
- , ,
, - “ ” , - ,
,
.
,
.
- , ,
, , ,
, ,
.
.
. , ,
, , ,
, , , ,
2009
65
.
(5- 11- )
, ,
,
.
.
,
.
2003
2-3 .
, ,
, .
, ,
,
.
3719 7 . . 3707
, 6 . . 343 ,
3 . . 3397 ,
. . 3526 .
, , , ,
, - “ ” “ ”
.
, ,
, .
,
, .
2009
66
- :
- ,
.
. 624,22 2 .
.
- 190 2 .
, -
.
. 60
.
- 638.70 2
2 .
.
- - - ,
136.9 2
,
- ,
,
.
- , 393,59 2 .
- 316.82 2
.
,
. ,
,
, ,
.
.
2009
67
18- . -
2001
-
, , , 670 2 40
, , , ,
650 2 20 , ,
2470 2 90 , ( ) , ,
780 2 200 , 6364 2 380
.
3164 2 380
, 1000 - 0.6 .
-
. , . 2007
2009
68
, , , ,
, 57 .
2009 ,
, - “ ” , , ,
, 16 83
(11- ).
11- . -
-
2 14 10 4 3 13 4 7 2
3 9 5 4 , 2 7 3 2 2
3 14 4 4 2 3 1 3 26 5 13 8
16 83 31 26 14 11
2 14, 3 13,
3 9, 2 7, - -
3 14, 3 26
.
258.28 2 14,
2057.57 2 13, , 597.59 2
9, 1228.27 2 7, - -
820.97 2 14, 258.28 2 26
.
12- .
2007 2009
- 14
- 13
- 9
- 7
- - 7 14
25 26
- 25 -
57 83
2009
69
, , , ,
, .
- , 2007 7 2009
14 . 2007 25
2009 26 .
,
.
, .
14 - . ,
2001
1:1.4, 1:1.5, 1:1.7 2007
: : : 2.3: 4.8: 1: 1.1 . :
1:2.07, 1:4.8, 1:4.1
.
3 (21.4%)
, (35.7%) , 6 (42.9%) .
,
1: 5.8 .
-
2001 -
6364 2 380 .
2009
70
-
. , . 2007 -
4.597 484±24
. 2001
27.3%- .
1-13- ( x =5.2, SD = 3.4). 83
, 38 , 20 , 16 , 9 .
(> 2 ) ,
.
2 (hazard rate key function with a maximum of 2
cosine adjustments, half-normal key function with maximum of 2 cosine adjustments)
, .
1,236 (SE = 282m)
0.321 / 2 (95% 0.137 – 0.753),
12,162 2 1550 .
2009 , 2001
.
-
.
3900 .
.
.
- 6449,7 ² .
2 11 11 50
.
- multiply survey 4 -
25,2
² (S=2 R) .
207,2 ² 50 .
6449,7 ² [=6449,7*50/(2*3,14*4*11)] 1550±6
.
2009
71
-
1:500,000
, .
. - 32
314 . 7 34
. 12
140
. , 13 314
258-
135 , 71 , 34 , 18 .
(> 2 ) ,
.
, ,
, . , -
, , , -
, ,
.
6 78
. 14
.
:
, , , , ,
, , , , , ,
, , , -
, , , , ,
, , , , ,
, , , , , ,
, , , , ,
, , , , , ,
, , , ,
, , , , ,
, , , ,
, , - , , ,
, , , , , ,
2009
72
, , , , , , -
, , , , , ,
, , .
2006
281 10 2 12,54
.
, , ,
, , , ,
, , ,
, , 1155,6 2,
, , , , , , ,
, , , ,
, , 2 , ,
, , , , ,
, , ,
.
19- .
2009
73
- “ ”
.
. , ,
. , , ,
, .
100 .
-
,
, .
, ,
. ,
,
. 8 10
.
2007
, ,
.
, ,
.
13- . -
2007 2009 - 71
- 112 - 3
- 14 - 14
- 7 - - 14
- 30 197 47
- 16 4 -
11 - 26 -
2 - 1 -
241 328
2009
74
, ,
,
,
, .
14- . ,
-
2009 32 328 129 77 38 17 54
2007 6 241 23 99 90 29
2007 241 23 (9.5%) , 99 (41%)
, 90 (37.3) , 29 (12.2%) 2009
32 328 , 129 (39.3%),
77 (23.4%), 38 (11.5%), 17 (5.1%) .
2007 2009 ,
. , 2007
2009 16 .
, .
, , ,
.
,
.
-
-
. , . -
21543,07 2 5720±286
1000 - 2.6 .
13 154
.
1,865m, (SE = 92m)
0.404 / 2 (95%
0.195 – 0.837), 12,162 2 3
2009
75
4913 (95%- 2,372 - 10,180)
.
.
10 , , , , ,
, , .
- ,
.
.
. , (16
83 )
.
,
. -
, . -
, .
.
.
15000
.
.
.
. 2001
6 3142,38 2
, , - ,
,
. 2009 ,
, - , ,
,
.
2009
76
, ,
(2001 ). ,
. 2
, .
.
.
.
,
.
,
(2001 ) .
2008
,
. -
,
. -
2009 10- 10- 2010
5-
.
2009
77
20- .
- , ,
(2001 ) .
, -
. ,
. .
,
(2001 ) .
.
2009
78
(2001 ).
.
,
(2001 )
, .
(20- ).
,
1<5 6 ( 40,0%)
18 ( 12,6%), 6<10
5 ( 33,3%) 42 (
29,4%), 11<15 2 ( 13,3%)
27 ( 18,9%), 16<20 1 (
6,7%) 16 (
11,2%), 31<40 1 ( 6,7%) 40
( 28,0%) (15- ) .
15- .
, 100:62.5, , 100:6.3, ,
100:34.4 .
6.3% 34.4% .
, ,
15 143 14
62 . 20
2009
79
14.0% , 32 22.4% , 2 1.4%
, 11 7.7% . 12
4,393 ² 3,561 ²
50 95%-
444-927,
572 .
2009
49,3%- .
.
( ) .
. ,
, , , .
, ,
,
, .
.
. ( ) 6
39 310 .
1- 50- ( x =7.9, SD = 9.1). 192 ,
90 , 53 , 1 , 48 .
(> 2 ) ,
. , ,
1,160 -2,368, 4
1,756 . , .
1000 .
, 50-60
2009
80
, .
.
, 50-60
. , ,
, ,
.
, ,
50-60, 10-20, 10 ,
. 2-3
,
. ,
. 2
10 . 21-
.
21- .
4 (2 , 2 ) . ,
, , , ,
2009
81
.
, (22- ).
2568.9 2 330
100 - 1.3 .
22- .
3 12 , 2 ,
2 .
.
31 23-
8,743 2 80.8% . 194 (
1-41). 112 .
17 102 . 23 11-
. 1- 22- ( x = 4.6, SD = 4.7).
2009
82
66 , 41 , 15 , 3 , 7
. (> 2 )
, .
half-
normal key function with maximum of 2 cosine adjustments
1,006 (SE = 140 ).
0.275 / 2 (95% 0.137 – 0.555),
2,404 2 2,123 (95% 1,198 - 4,852)
.
2001
14 9937 2 3044
6.1% , , , , 4.8%
, , , , 3.2%
, , , .
- 473 2 60 ,
, 316 2 50 , - ,
481 2 140 , 570 2 110 , , 348 2
90 , , 86,0 2
50 , , , ,
1792 2 1150 , - 42 2 20 , , , 1395 2 280 , , , 1040 2
460 , 629 2
210 ( 2001).
2007
8890.77 2 956
, 1000 - 1.06
.
4945.8 2 2018
, 1000 - 4.08
( 2007).
2009
83
10- , 683 2
.
2007 100 , 553 2008
48 192 , 108 743
.
-
.
- , , ,
, , ,
, , , ,
, , , ,
251 (
) . ,
11 175 , 77
. ,
11 25 , 24 ,
, , .
,
.
G-10 9 32 , 23
. -
,
. ,
.
6 5.3
28 12 , 3 , 5 , 8
. 43 %- ,
1.5:1
.
2009
84
4 23 5.75
.
.
. 9 35 , .
6 5.83
, .
20 11 , 3 , 2 , 4 .
55 , 15 , 10 , 20
. ,
. 3.66:1
.
9
2008
, – , –
. ,
.
, 21 44 , 9 .
.
. - ,
.
.
, ,
.
. 9
4.88 .
.
33 13 , 8 , 7 , 5 .
2009
85
40 , 24 , 21
, 15 . 2.66:1
.
1.66:1 .
3 .
, .
. 3 .
12 12 , 1 .
2007 , ,
. 3
4 . 2 8
.
, ,
. 24
29 , 2 .
, , ,
,
.
.
13 2.2
1 . 26 11
, 3 , 8 , 4 . 42
, 12 , 31 , 15 .
2.8:1
. 3.5:1 .
9 2 , .
21
, 7 , ,
.
.
2009
86
. , .
3 ,
, , “ ”, “ ”
.
( ) 10
14 .
. 6
2.5 8 3 , 1 , 2 ,
2 . 1.48:1 ,
2.8: 1 2.8 .
.
3 14 ,
9
. 3 2 , 6 , 1 , 2 .
2 . ,
.
. 2007 3
. 10
.
. 2
.
14 3 5 .
.
.
. . .
17 22 .
, ,
, .
.
2009
87
7 25 .
.
23- .
. , ,
.
2001, 2007
,
. 2001 9937 2
4044
.
25 14 56% , 7 28%
, 4 16% .
, , ,
.
,
, (568,48 2), (113,65 2) . ,
, , ,
. 5 24
.
2009
88
24 4 17% , 11 46% , 1
4% , 6 25% 2 8%-
.
.
24- .
,
1990- . , .
1995 1000 - 5,7 , 0,8 , 1996
1000 - 6,4 , 1,2 .
1995 2977, 1996 3333 , 1997
3250±1071 3000-3300
(Reading R.P 1997).
2001
14 9937 2 3044
1000 - 3.1 ( . 2001).
2007
8890.77 2 956
1000 - 1.06
. 4945.8 2 2018
, 1000 - 4.08
( 2007).
2009
89
31 23-
8,743 2 80.8% . 194 (
1-41). 112 .
17 102 . 23 11-
. 1-22- ( x = 4.6, SD = 4.7). 66
, 41 , 15 , 3 , 7
. (> 2 )
, .
.
half-normal key function with maximum of 2 cosine adjustments
1,006 (SE
= 140 ). 0.275 / 2 (95% 0.137 –
0.555), 2404 (95% 1,198 - 4,852)
.
13 6
. 249 109- , 78 , 6 , 2
, 23 . 52 (190 ) distance
. 2 (hazard rate key function
with a maximum of 2 cosine adjustments, half-normal key function with maximum of 2
cosine adjustments) 530
(SE = 682 ). 1.524 / 2 (95%
0.877 – 2.65), 12,162 2
13324 (95% CI 7,671 - 23,169)
.
2009
90
16- .
2000- 3000
2001 2007
. , ,
2000
. 2009 2404
, . 2007
( , , , )
.
,
,
.
.
, . ,
.
, .
2009
91
,
.
( X = 530 )
.
,
,
.
,
.
. ,
.
, .
.
, .
, , , ,
.
200 1 2009 10- 15-
. - 10 12- 7
.
2009
92
.
1401 2 24 21-
11 .
25- .
72 . 19 142 .
1-17- ( x = 7.5, SD =4.6). 142 ,
84 , 26 , 7 , 25 .
half-normal key function
with maximum of 2 cosine adjustments
894 (SE = 1160 ).
0.595 / 2 (95% 0.298 – 1.188), 1,401 2 834
(95% 417 - 1,664) .
1
11 ( ) .
, .
2009
93
26- .
2009 10- 6- 57 , 10-
8- 11 .
, , .
.
0.6 / 2 .
, .
, .
.
11 4,010.05 2
8 3,587.6 2 , 6
494.22 2 , ,
. 2009 11 . - ,
, , , ,
2009
94
,
.
- - , , ,
, , , ,
,
.
27- .
46 293
20 (8%) , 176 (75,%) , 3(1%) , 37 (16%)
. 2-12
2007 2009
0,6%- 1,2%, 11,2%, 1,8%- .
: : : 6,7:58,7:1:12,3 .
1,7% .
98,3 .
- 58,7:1
.
2009
95
, :
2007 1.6:1 2009 4.8:1 .
,
,
, , .
2001
, 410 2 100 ,
186 2 20 , ,
410 2 110 , 886 2
10 , , , , ,
, , 957 2 200 ,
, 421 2 60 ,
342 2 20 , - 155 2
30 , 157 2 20 ,
, 126 2 110
, 3924 2 850 .
21 18- (
4,439 km2 96%). 108 6 . 90
( 2-9) . 49
. 46 293 . 1-27-
( x = 6.2, SD =5.9). 236 ,
176 , 20 , 3 , 37 .
half-normal
key function with maximum of 2 cosine adjustments
619 (SE = 1370 ).
1.084 / 2 (95% 0.339 – 3.471), 4,439 2
2330 .
2007 2001 1,5%-
2009 2007 0,7%- .
2007 2001 1,3%- , 2009 2007 30,6%-
.
2009
96
.
4800
.
.
.
1723,5 ² .
46 58 293
.
49 multiply survey
.
line transect 108
, 2 -
216 ² .
216 ² 293 .
1723,5 ² [=1723,5*293/216] 2338
.
5 1564.7 2
.
, ,
, , ,
.
2009
97
28- . , 2009
,
. ,
, - , ,
, , ,
14 75 .
.
7
,
10 (28,6%) , 16 (45,7%) , 2 (5,7%) ,
32 (20%) .
: : : 5:8:1:3,2 . :
8:1 . :
8:3,2 43,8%- . , ,
,
. 2007 2665.4 2
. 2009 21 6- .
1-20- ( x = 5.3, SD =5.1). 35 ,
16 , 10 , 2 , 7 .
2009
98
half-normal key function with maximum
of 2 cosine adjustments
703 (SE = 65 ).
0.634 / 2 (95%
0.2889 – 1.392), 4439 2 1520 (95%
1,278 - 6,179) .
. :
1.
2. ,
,
3. ,
,
4. ,
5.
.
5 (310, 75, 6, 5,35) , 1 6 , 4 , 1
.
Distance sampling
, .
,
. ,
K.
.
, , (40-50 ),
, (10- ), (10
), (60-70 ), , , (3-4),
2009
99
(15 ), , , , (100
), (10), , , (20 ),
, , , , , ,
, , , , , ,
, , , ,
, , , , ,
, , , , , , ,
, , , , , ,
9825,4 2 (29- ).
, , ,
. , ,
. .
. ,
. .
,
.
1-32
. 1-5 (14 ) 41,2%-
. 1-10 70,6%- . 15-
, 17- .
15- .
% % 1- 5 14 36 41.2 11.0 6- 10 10 80 29.4 24.4 11- 15 2 24 5.9 7.3 16-20 3 55 8.8 16.8 21-25 2 44 5.9 13.4 25-30 2 57 5.9 17.4 31-35 1 32 2.9 9.8
2009
100
17- .
0.05.0
10.015.020.025.030.035.040.045.0
1- 5 6- 10 11- 15 16-20 21-25 25-30 31-35
% %
, 1- 22
(..- , ..- ). 1-5 (49)
69,0%- . 1-10 87,3%-
63,8% .
16- .
% % 1- 5 49 143 69.01 38.34 6- 10 13 95 18.31 25.47 11- 15 6 78 8.45 20.91 16-20 2 35 2.82 9.38 21-25 1 22 1.41 5.90
71 373 100.00 100.00 14.20 74.60 20.00 20.00
SE 8.95 21.73 12.61 5.82
18- .
2009
101
, .
, , 34 328
. 282
86.6% .
: : : 23,4: 69,9: 3,2: 3,5 .
67 .
, 841 -3,561, 4
2050 .
.
. 324 .
~ 121 32
189 . DISTANCE 1.63
/ 2 (95% 0.59 – 3.29) , 539 (95%
196-1,082) .
– 467 .
24-98, 58.1 . 4
19-223-
91.4 . 25 8
(7 , 1 ) ( 12 ). 7 .
461 (95%- 278 – 643), 191 (95%-
81 – 379), 652 (%- 359-1,022) 779
(95%- 269 – 1,289), 268 (95%-
5 – 820), 1,047 (%- 274 – 2,109)
.
( ) 2773 .
2009
102
29- .
,
, , -
.
- 2009 9, 10, 11-
.
98 941 . 1-5
35,7%- , 1-10 66,39-
. 10-
.
2009
103
16- . -
% % 1- 5 35 119 35.71 12.65 6- 10 30 230 30.61 24.44 11- 15 15 194 15.31 20.62 16-20 9 161 9.18 17.11 21-25 5 109 5.10 11.58 26-30 2 55 2 6 31-35 0 0 0 0 36-40 2 73 2 8
98 941 100 100 12.25 117.63 12.50 12.50
SE 4.75 26.78 4.85 2.85
19- .
6 22
, .
,
. -
2000 .
2009 9, 10, 11-
. 3
: : : 26,9: 41,8:
12,6: 18,7 (20- ).
2009
104
20- . -
100 ,
10 ,
10 , 120
.
30- .
2009
105
2008 12-
.
.
- .
.
, -
“ ”
10 .
.
, .
, , ,
, , , , ,
GPS, .
,
, ,
.
, ,
. 2000 ,
2005
.
. 10
3 1000- .
2 ,
.
2009
106
.
.
.
. , New Juulchin tour
1 .
( ) .
2005
( - ) 4
.
.
.
. -
.
.
-
.
.
.
, , , , ,
6 , 2 .
. .
1976 , 2 100
- . ,
. . ,
2009
107
19 140 . 139
. : : :
18,7:56,8:2,9:21,6 .
17- .
100 5,1 38,0
.
.
.
.
.
. .
-
.
2009
108
, , , ,
14 682,5 2 180
.
31- .
.
3 135 5 , 23
. , .
. ,
27 . 7
, 40
.
.
1389 2 36-61 2
.
.
2009
109
32- .
3 22 .
100 - 7
1335 2 1330
.
33- . .
2009
110
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