ACADEMIE POLONAISE DES SCIENCES

INSTITUT DE PALEOZOOLOGIE

PALAEONTOLOGIA POLONICA - No. 20,1968

FORAlVIINIFERA FROlVI THE PALEOCENEOF POLAND, THEIR ECOLOGICAL

A.ND BIOSTRATIGRAPEIICAL MEANING

(OTWORNICE Z PALEOCENU POLSKI, ICH EKOLOGICZNEI BIOSTRATYGRAFICZNE ZNACZENIE)

BY

KRYSTYNA POZARYSKA & JANINA SZCZECHURA

(WITH 22 TEXT-FIGURES, 3 TABLES AND 18 PLATES)

WARSZAWA 1968

PANSTWOWE WYDAWNICTWO NAUKOWE

R EDAKTOR - R ED AC TE UR

ROMAN KOZLOWSKICzl onek rzeczywisty Po lskie] Akademii Na ukM embre de I'Acadernie Pol on a ise des Scienc es

Redaktor techniczny - R edac teur tec h niq ue

Wcro nika Sipo wicz

Adr cs Rcd akc] i - Ad resse de la Red uctio nInst itut de Pal l:o zoo logic

de l'Acadcmie Po lon aise des Sci encesWa rszawa 22. A I. Zwir k i i W igur y Nr 93

Co pyr igh tby Panst wowe Wydawnictwo Nau kowc

1968

Prin ted In Poland

Pa nstwo we W yd awnict wo N auk o we - Warszawa

N akIad 650+90 egz, Ar k. wyd. 13,5.Arkuszy d ruk, 6"/.. + 19 wkladek ,Papicr dr uk, sa t. k l. 111 64 x 96 100 g,Odda no do sk lada nia 12. VII . 1967 r,Podpi sano do d ruk u 30 . X . 1968 r.Dr uk ukon czon o w l istopadz ie 1968 r.

D rukarnia U niwcrsytetu J agiclloriskiego w Krako wioZam. 832167

CONTENTS

Preface . . . . . . .Ackn owledgements

GENE RAL PART

The lowermost Tert iar y in the Pamietowo bor ing , North Pol and . . . . . . . .Th e Boreal Province in Europe and th e region s with simi la r biofacies co nd itions

Den mark and Sweden .Austria .England .U.S.S.R. (European and As iatic pa rt)

Lithuan ia .Dnieper-Donetz Bas in .Ukraine .Crimea .Ge orgia .Tadzhik depression

North America . . . .Au stralia. . . . . . .

The Meridional (non Mediterranean) ProvinceFrance (Paris Basin) . . . . . . . . .SW Belgium (Mons Basin) . . . . . .NE Belg ium and S Holland (Limburg Province)Germany .

The stra t igraphy of the lowermost Tertiary in EuropeConclusions . . . . . . . . . . . . . . . . . . . .

SYSTEMATIC PART

Family Textulariidae EHRENBERG, 1838. . . . . . . . . . . .Genus Splroplectammlna CuSHMAN, 1927 .

Sp iroplectammina wilcoxensis CUSHMAN & PONTON, 1932Genus Textularia D EFRANCE, in DE BLAINVILLE, 1824 .

Textularia plummerae arkansasana CuSHMAN, 1951Family Ataxophragmiidae SCHWAGER, 1877

Genus Arenobulimina CUSHMAN, 1927 .. .Arenobulimina cusk leyae JENNINGS, 1936

Genus Vacuovalvulina HOFKER, 1966 . .Vacuovalvulina keijzeri (VAN BELLEN, 1946)

Family Lagenidae CARPENTER, 1862 . . .Genus Astacolus DE MONTFORT, 1808 .

Astacolus arcuatus (PHlLIPPI, 1843)Astacolus gryi BROTZEN, 1948 . . .Astacolus paleocenicus BRoTz EN, 1948

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Genus Frondicularia D EFRANCE, in O'ORBlGNY, 1826Frondicularia bifor mis MARSSON, 1878

G enus Marginulina O'ORBIGNY, 1826 .Marginulina costulata HOFKER, 1958

Genus Robulus DE MONTFORT, 1808R obulus degolyeri (PLUMMER, 1926)Robulus rancocasensis OLSSON, 1960R obulus turbinatus (PLUMMER, 1926)

Family Pol ymorphinid ae O'OR BlGNY, 1839Genus Globulina O'ORBlGNY, in OE LA SAGRA, 1839

Globulina arenacea BROTZEN, 1948. . .Globulina gibba gibba (O'ORBIGNY, 1826) .Globulina g ibba multistr iata n . subsp . . . .Globulina cf. luberculata O'ORBlGNY, 1846Globulina sp . . . . . . . . . . . . . . .

G enu s Guttulina O'ORmGNY, in DE LA SAGRA, 1839Guttulina communis O'ORBlGNY, 1826Guttulina problema O'ORBIGNY, 1826 . . .

G enus Pseudopolymorphina C USHMAN & OZAWA, 1928Pseudopolym orphina fronde a (CUSHMAN, 1922)Pseudopolymorphina geijeri angusta BROTZEN, 1948Pseudopolym orphina paleocenica BROTZEN, 1948

Genus Pyrulina O'O RBlGNY, in OE LA SAGRA, 1839Pyrulina fusijormis (ROEMER, 1838)

Ge nus S igmomorphina CUSHMAN & OZAWA, 1928 .Sigmom orphina brotzeni HOFKER, 1957 . . . .S igmomorphina pseudo regularis C USHMAN & THOMAS, 1930

Family Gl andulinidae R EUSS, 1860 .G enus Glandulin a O'ORBlGNY, in OE LA SAGRA, 1839

Glandulina laevigata O'ORBlGNY, 1826 . . .Family Turr ilin idae C USHMAN, 1927 . . . . . . . . .

G enu s Pyramidina BROTZEN, 1940, nom . nu d. , emend. BROTZEN, 1948Pyramidina crassa BROTZEN, 1948 .Pyramidina cuneata (BROTZEN, 1948)

Fa mily Boli vinitidae CUSHMAN, 1927 .Genus Bolivina O'ORBlGNY, 1839 .

Bolivina oedumi BROTZEN, 1948Family Buliminidae lONES, 1875 . . .

Genus Bulimina O'ORBIGNY, 1846 .Bulimina ovata O'ORBlGNY, 1846 .Bufimina paleocenica BROTZEN, 1948

Family U vigerinidae H AECKEL, 1894 . ..G enus A ngulogerina CUSHMAN, 1927

Angulogerina europaea CUSHMAN & EOWAROS, 1937Family D iscorbidae EHRENBERG, 1838 . .

Genus Discorbis LAMARCK, 1804 . . . . . . . .Discorbis quadrata (TERQuEM, 1882) . . . . .

G enus Epistominella HUSEZIMA & M ARUHASI, 1944Epistominella cf. limburgensis (VISSER, 1951)

Genus Rosafina O'O RBIGNY, 1826 . .R osafina brotzeni HOFKER, 1961 .Rosalina koeneni BROTZEN, 1948 .Rosalina selandiana n. sp.Rosalina ys tadiensis BROTZEN, 1948

Genus Rotorbinella BANOY, 1944

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Rotorbinella mariei (VAN BELLEN, 1946)

Rotorbinella mont iana n. sp,Rotorbinella papillata n. sp. . .

Famil y Spirill inidae R EUSS, 1862

Genus Patellina WILLlAMSON, 1858

Patellina sp. . . . . . . . . .

Ge nus Spirillina EHRENBERG, 1843

Spirillina cr. nodifera T ERQUEM, 1882

Fa mily R ota1iidae EHRENBERG, 1839 .

G enu s Cuvil lierina D EBo uRLE, 1955 .

Cuvillierina ? pomeraniana n. sp. .

G enu s Pararotalia L E CALVEZ, 1949 .

Pararota/ia tuberculifera (REUSS, 1862)

Genus Rotalia LAMARCK, 1804 . . . .

Rota/ia marginata D'ORllIGNY, 1826

Rota/ia saxorum [)'ORBI GNY, 1850 .

Rotalia trochidiformis (LAMARcK , 1804)

Family Elphid ii dae GALLOWAY, 1933

Genus Elphidium DE M ONTFORT, 180S .. .

Elph idium cr. lama rcki (D' ORBlGNY, 1826) .

Genus Elphidiella CUSHMAN, 1936

Elphidiella prima (TEN D AM, 1944) .

Fa mi ly Glob orota1i id ae CUSHMAN, 1927

G enus G/oboro ta/ia C USHMAN, 1927 . .

G/oborotalia globigeriniformis VAN BELLEN, 1946 .

G/oboro talia praepseudom enardii H OFKER, 1961

G/oborotalia sp. . . . . . . . .

Ge nus Globorotalites BROTZEN, 1942

Globorotalites gra nulatus n. sp.Family Globiger inidae CARPENTER, PARKER & JONES, 1862

G enu s Globigerina D' ORllIGNY, 1826 . .. . .

Globigerina aff. compressa PLUMMER, 1926

Globigerina daubjergensis BRONNIMANN, 1953

Globigerina k ozlowsk ii BROTZEN & POZARYSKA, 1961

Globigerina pseudobulloides PLUMMER, 1926 . . .. .

Globigerina (Subbotina) triloculinoides PLUMMER, 1926

Globigerina trivialis SUBllOTINA, 1953 .

Globigerina varianta SUBBOTlNA, 1953

Family Eponididae HOFKER, 1957 .. .

Genus Eponides DE MO NTFORT, 1808

Eponides toulmini BROTZEN, 1948

Genus Neoeponides R EISS, 1960

Neoepo nides cf. schreibersi (D' ORBlGNY, 1846) .

Fami ly Cibicididae CUSHMAN, 1927

G enu s Cibicides DE M ONTFORT, 1808 .

Cibicides asteroides n. sp. . . . . .

Cibicides aurou zae ROUVILLOIS, 1960

Cibicides carinatus (TERQUEM, 1882)

Cibicides cuvillieri ROUVILLOIS, 1960

Cibicides lectus VASILENKO, 1950

Cibicides proprius (BROTZEN, 1948)

Cibicides sahlstr oemi BROTZEN, 1948

Cibicides simplex BROTZEN, 1948

Cibicides succedens BROTZEN, 1948

Genus Planulina O'ORBlGNY, 1826

5

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Planulina /imbata BROTZEN, 1948 .Family Nonionidae SCHULTZE, 1854 . .

Genu s Nonion DE MONTFORT, 1808 .Nonion graniferum (TERQUEM, 1882)Nonio n sublaeve TEN DAM, 1944 . .

Genus Nonionel/a CUSHMAN, 1926.. .Nonionel/a communis communis (D'ORBIGNY, 1846).Nonionel/a communis paleocenica n, subsp.Nonionel/a ovata BROTZEN, 1948 .

Family Alabaminidae HOFKER, 1951 . . . ..Genus A labamina TOULMlN, 1941 . . . . .

Alabamina midwayen sis BROTZEN, 1948 .Family Osangulariidae LOEBLlCH & TAPPAN, 1964 .

Genus Gyroidinoides BROTZEN, 1942 . . . . .Gyroidinoides octocamerata CUSHMAN & HANNA, 1927Gyroidinoides pontoni BROTZEN, 1948 . . . . . . . .

Genus Osangularia BROTZEN, 1940 .Osangufaria cordieriana navarroana (CUSHMAN, 1938)

Fami ly Anomalinidae CUSHMAN, 1927 . . . . .Genus Anomalina D'ORBIGNY, 1826 . .

Anoma/ina burlingtonensis (JENNINGS, 1936)Anoma/ina danica (BROTZEN, 1940)Anomalina ekblomi (BROTZEN, 1948) .Anomalina min or n. sp.. . . . . . .Anoma/ina praeacuta VASILENKO, 1950Anomalina umbi/icata costata n. sub sp..Anoma/ina umbi/icata umbi/icata (BROTZEN, 1948)

Genu s Coleites PLUMMER, 1934 . . . .Coleites reticulosus (PLUMMER, 1926) .

Genu s Gave/ine/la BROTZEN, 1942 .Gaveline/la le//ingensis BROTZEN, 1948

Genus Karreria RZEHAK, 1891 ..Karreria falla x RZEH AK, 1891 .

Genus Pulsiphonina BROTZEN, 1948Pulsiphonina prima (PWMMER, J926)

Family Ceratobuliminidae CUSHMAN, 1927 .Genus Ceratobu/imina TOULA s.s ., emend. FINLAY, 1939 and TROELSEN, 1954 .

Ceratobulimina tuberculata BROTZEN, 1948Genu s Lamarckina BERTHELlN, 1881

Lamarckina rugulosa PLUMMER, 1926 .Genu s Mississippina HOWE, 1930 . . . .

Mississippina binkhorsti (REUSS, 1862)Genus Stomatorbina DORREEN, 1948.

Stomatorbina sp• .Bibliography. . . .Alphabetical indices . .

Index of authors . .Palaeontological index '.

Plates I-XVTII and their legends.

Page808181818182828383848484858585858585868686868787888989909091919191929293939393939494949496

102102104

PREFACE

The pr esent paper is a supplement to existing publications on lowermost Terti ary inPoland (POZARYSKI & POZARYSKA, 1960; BROTZEN& POZARYSKA, 1957, 1961) and in particularto POZARYSKA'S monograph (1965) on Dano-Montian Foraminifera of Central and NorthernPoland. It is ba sed on investigations of the Parnietowo boring (North Poland) , where the presentauthors ob served a mergi ng of the influence of borcal an d meri dional (non Mediterranean)Paleocene. T he superposition of boreal and mer idional Pa leocene format ion s in Poland and theana lysis of Paleocene formations outside Po land allowed the authors to correlate the lower­mo st Tert iary beds in Europ e and to a certai n extent those beyond it. The authors came to,among others, the conclusion that the Mo nti an s. s. is part ly of th e same age as the Selandians. s. However, at the ma rgina l area of Boreal and Mer idiona l Provinces, i. e. mainly in Poland,the Paleocene subtropica l sediments cover th ose exclusively containing a boreal microfauna.The Paleocene is here con sidered as the oldest stage of Tertiary embracing Danian, Montianand Thanet ian. Mo ntian s.s. an d Selandia n s.s. are con sidered as partly stratigraphica l­facies un its. Th e term Selandian was in 1924 in troduced by ROSENKRANTZ to include the Pa­leocene ser ies of strata which in Den mark and Scan ia is rest ing on the D anian and is over­laid by the Lower Eocene ash layer series. Thi s div ision of the lowerm ost Tertiary is acceptedin the present pap er.

In the present pa per, 97 specie s (belonging to 50 genera an d 21 families) are described,out of which 7 species and 2 subspecies are new ones . Their systematics is taken from theTreati se on In vert ebra te Paleontology (by LOEBLlCH & TAPPAN), C, 2, 1964, and is not revisedhere. The main atte ntion is concentrated on the descripti on, d istribution and variation of thedetermined spec ies, the variatio n, however, being treated in a rather wide sense. The citednumber of spec imen s for each given species includes specimens no t only fro m the Parnietowoboring, bu t also those fro m the Paleocene of Central Poland, i. e. those examined earlier byPOZARYSKA (I965).

The collection of specimens figured in th is paper is ho used in the Palaeo zoological In­stitute of the Poli sh Aca demy of Sciences, for which the abbrevia tio n Z. Pal. is used.

ACKNOWLEDGEMENTS

The investigated mater ial was obtained from the Geological Institute in Warsaw, to thea uthorit ies of which the aut hors wo uld like to exp ress their grati tude .

The present a uthors a re greatly ind ebted to Professor ROMAN KOZLOWSKI for his in­valua ble help, ad vices and discussions during the present investigations.

Sincere than ks are due to the following a uth ors for kindly send ing comparative materialas well as for their valua ble discussion and rem arks: Prof. H. M. BOLL! (Geol. Inst., Eidg.Techn. Hochschule, ZUrich), the late Dr. F . BROTZEN (Swedish Geol . Survey, Stockholm),

8 KRYSTYNA POZARYSKA & JANINA SZCZECHURA

Dr. Y. LE CALVEZ (Lab. de Micropaleontologie, Ecole Prat. des Hautes Etudes, Paris),Mr. D. CURRY (London), Prof. J. CUVILLIER and Dr. A. ROUVILLOIS (Lab. de Micropaleon­tologie, Universite, Paris), Dr. J . HAYNES (Dept. of Geology, University, Aberystwyth),Dr. J. HOFKER, Sr. (La Haye), Dr. P. MARIE (B. R. G. M., Paris), Prof. R. MARLIERE (Dept.of Geology, Polytechnic School, Mons), Prof. M. M. MOSKVIN (Dept. of Geology, University,Moscow), and Prof. A. ROSENKRANTZ (Dept. of Geology, University , Copenhagen).

Sincere thanks are also due to Mrs. E. G ADOMSKA for the preparation of drawings,plates and text-figures, Mrs. K. BUDZYNSKA for a map, and Mi ss L. LUSZCZEWSKA forphotographs.

Palaeozoological Instituteof the Polish Academy of ScIences

Warszawa, AI. Zwirki i Wigury, 93April 1967

GENER AL PART

THE LOWERMOST TERTIARY IN THE PAMI~TOWO BORI NG,NORTH POLAND

(Text-figs. I , 2; Ta ble I)

The starting point of the present paper is the Parn ietowo boring near Chojnice in Po­merania, North Poland. This boring was dr illed in 1959 by the Geological Institute of Poland.The boring passed through a rather thin series of Quatern ary and Tertiary sediments, whoseage could not be precisely determined because of a complete lack of fauna, and then penetr ateda glauconitic-marly series yielding a considera ble amo unts of coa rse calcareous sand at thebase of the Terti ary beds. On examination this calca reous sand was found to be the same typeof sediment as the so-called "tuffeau" , represe nting ca lcareo us detritic sedime nt for med bycrushed , worn out, friable , calcareous, mainly organic sediment, con ta ining a great ad mixtureof glauconite. Th is was the first time, the so-called "t uffeau" type of sediment had been foundin Poland. Previou sly, it was known in series of that age in Belgium , Holland and France onone side, and in the Crimea on the other. In this detr itic series, which is about 70 m in thicknessin the Pamietowo boring, there occur several thin layers of hard limestone, interbedded withthe "tuffeau". Throughout the whole of this series there are rather small, gray concretionsof silica (chert s), although these are not very numerous. At the base of the series, smallphosphatic nodules occur as well as abundant remains of bryozoan colonies. In the Parn ietowoboring all thi s "tuffeau" series lies directly on a ha rd gro und layer, ending the Lower Maastricht­ian sedimentation. Th is latter layer is composed of white-gray limestone , containing a typicalforaminiferal assemblage with Rugoglobigerina rugosa (PLUM.), Stensiiiina pom merana BROTZ.,Bolivina incrassata (R EUSS) and others. Upper Maastrichtian and Danian beds are absent.

Macrofaunal remain s in the above mentioned series of "tuffeau" are sca rce, badly pre­served, almo st unidentifiable . They mainly consist of small pelecypod valves, worm tubes,single loose plates of echinids, spicules of silica sponges and othe rs. In contras t to the scarce,badly preserved macrofauna in the "tuffeau" series, there is a very rich , well preserved as­semblage of microfauna, thi s latter, comprising Foram inifera, being subject of the presentstudy.

The age of the "tuffeau" series in Pamietowo was determined, in a preliminary note ofBROTZEN and POZARYSKA (1961). It was there assigned to the Paleocene with the uppermost15-16 metres, erroneously considered as belonging to the lowermost Eocene. Determinationof the age, especially the lower part of the examined profile, was based, at that time, on anassemblage of Foraminifera analogous to that known and described from Sweden (BROTZEN,

10 KR YSTYNA POZAR YSKA & .IANI NA SZCZECHURA

1948) and Central Poland (BRoTzEN & POZARYSKA, 1957). The "tuffeau" series of Pamietowoyields typical Selandian Foraminifera, i.e. Scandinavian Lower Paleocene (ROSENKRANTZ,1924). In addition to the Selandian assemblage, there are in the Pamietowo boring, espe­cially in its upper part, several species previously unknown in the Lower Paleocene (Selan-

£ boring

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dian). These latter were provisionally considered, at that time (1961), as representing newspecies . Some of them were identified, however, as being very similar to those occurring in theBunde Beds in Holland, described by VAN BELLEN (1946) and referred by him to Eocene.

For further investigation of the Parnietowo boring it was necessary to obtain com­pa rative material. Several samples have been kindly sent by Prof. R. MARLIERE from Belgium,fro m the famous boring in Mons (Puits Artesien at Polytechnic Faculty), drilled a hundred

FORAMINI FERA FROM THE PALEOC EN E OF POLAND 11

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R. papil/ata n. sp.

Rotortunetla montian a n.sp.

Nonion subt aev e ten Dam

Cuviil ieriru: oomeraniana n.sp.

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tamorckina ruqulosa Plummer

sareunetta teltinqensi« Brotzen

stoborotatites qranutatus n.sp.

ALabamina midwaljcnsis Brazen

Pulsiphonina prima (Plummer)

Ceratobulimina tutierculata Brotzen

Rosalina koeneni Brotzen

6. var i ont o Subbotina

GloborotaLia prceoseuaomenartiii HorheT

G. tritocutinouies PLummer

Non ion graniferum (Terquem)

6Lobigerina trivia lis Subbotina

6. aff. compressa PLummer

Spirillinti cf. noiiitera Terquem

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Fiz, 2. Pamietowo boring, North Poland.

12 KR YSTYNA PO ZAR YSKA & J A NI NA SZ CZEC H U RA

T A B LE

Dl STRIBUTlO:\ I ll" I ;Ol\\MI~I FU\ \ I ~ Til E MO:\T IAi\ OF THE I' AMlI~TOWO BORI~(;

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Textularia plummerae ark ansasana C USHMA N

Arenobulimina cuskleyae J ENN1NGS

Vacuovalvulina k eijze ri ( VAN fln.I.EN )

Astacolus arcuatus (PHILl PP I)

A . gryi B ROTZEI'

A. paleocenicus Bno rzsxFrondicularia biformis M ARSSOS

Marginulina costulata H OEKl- R

Robulus degolyeri ( P LUMM1R) .

R. rancocasensis O I.SSON

R. turbinatu s ( PUJMMER)

Glohulina arenacea B ROTH N

G. gibba gibba (I)'ORBI( ;NY)

G. gibba multist riata n. subsp.G. cf. tuberculata I) 'ORBI(''''

Globulina sp.Guttulina communis l) 'O R61<;'"

G. problema O' ORBIGS\

Pseudopolymorphina frondca (CUSI IM AN)

P. geijeri angusta B ROTZI '

P. paleocenica B RO I.ZEI'

Pyrulina fu siformis ( R OI.'II'R)

Sigm omorphin a brot zeni H orx ru

S. pseudoregularis C u smlA S & TI\O~I A S

Glandulina laevigata l) ' O RIl I( ;S\

Pyramidina crassa B 110 1ZL "

P. cuneate (BROI ZL N)

Bolivina oedum i BRo r Z EN

Bulimina Ol'ala O'ORRlG NY

B. paleoccnica BROT ZEN

Angulogerina europaea CUSII\I A'i & EI)W\RI"

Discorbis quadrata (T rnour v)Epistomin ella cf. limburgensis (V ISSI' R )

Rosalina brotzeni H OI.KER

R. ko eneni B ROTZ U '

R. selandiana n. sp. .R. vstadiensis Bnor zu-Rotorbinella mariei ( VA:" B I'I .L1·s) ,

R. mont iana n. sp,R. papillata n. sp.Patellina sp.Spirillina c f. nodifera TE RQUHI

Cuvillierina ? pomeraniana n. sp.Pararota lia tuberculifcra ( REUSS)

Rotalia ma rginata l) ' ORRlGNY

R. saxorum D' ORBIGN Y

R. trochidiformis ( L M tARCK)

Elphidium cf. lamarcki (D'ORBIGNY)

FORAMINIFERA FROM THE PAL EOC ENE OF POLA ND 13

TABL E

(continued]

_______ Depth (in Ill)

Species _____

I

-

i--

I 'I

I

I

+--+-I ,

-+-

,, ,-

I !'-, -I- -

I, ,:---:

,II

, ,,

L:::Ji II I

I

,I ,

~----t- ; !: :

t ; I

,

-,---'

!_ - ' ,; j J 1-r

r-I I : I I

: i

,

i I

_ I' I- - I1I

! I I~_ .- : I , - ,- --1--- i- :=""+----c::.- -

I- - ---' - +--t==---;-

i - - .- t----' - -

I- --+-- -;-- f-- -t - --:f-~----+--!---f---- ,-+----,----1-+--+--~"':--:""

Elphidiella prima ( TEl' D AM )

Globorotalia globigerinifor m is VAl' B ELLEN

G. praepseudomenardii H OFKER

Globorotalia sp.Globoro talites granulatus n. sp.Globigerina afT. comp ressa PLUMMIR

G. daub jergensis BR ONN IMA Nl<

G. kozlowsk ii BR OTZEl< & POi.ARYSK A

G. pseudohulloides PLUMM ER

G. {Subbotinu ) triloculinoides I'LUMM ER

G. trivialis S UBBOTl SA

G. varianta S UBBOTl NA

Eponides toulmini B ROTZ ES

Neoepo nides cr. schreibersi (O'ORllI ( ;S Y)

Cibicides asteroides n. sp.C. aurouzac R OUVI LLOIS

C. carinatus ( T ERQUEM)

C. cuvillieri R OUVI LLOIS

C. lectus VASILENKO

C. proprius ( BRO TZ El<)

C. sahlstro enti B ROTZEN

C. simp lex B ROTZ EN

C. succedens B ROTZES

Planulina limbata B ROTZEN

Nonion graniferum (TERQUEM)

N . sublaeve TEN D AM

Nonion ella commu nis communis (I>'O RBIGSY )

N . communis paleocenica n. subsp.N , ovata BR OTZ EN

Alabamina midwayensis B ROTZEN' ,

Gyroidinoides octo came rata CUSH MAN & HAN NA

G. pontoni B ROTZEN

Osangularia cordieriana navarroana (CUSH MAN) .

Anoma lina burlingtonensis (JI'NNINGS)

A. danica (BROTZEN)

A. ekblomi (BROTZEN)

A. min or n. sp.A. praeacuta V ASILENKO

A. umbilicata costata n. subsp ..A. umbilicata umbilicata ( BR OTZE N)

Coleites ret iculosus ( P LUM MER)

Gavelinella lellingensis B ROTZEN

Karreria falla x R ZEHA K

Pulsiphonina prima ( PLU~tMER)

Cerat obulim ina tuberculata B ROTZIoN

Lamarckina rugulosa P LlIM MER

Mississippina binkh orsti ( R EUSS)

St omatorbina sp.

14 KR YSTYNA POZAR YSKA & JANINA SZCZECHURA

years ago, where the true Montian, according to MARLlERE, was pierced and the Montian asa lower substage of Paleocene established. The present authors also had at their disposalsingle samples from the Bunde Beds (Limburg Province, Holland), as well as some samplesfrom the English Thanet Beds. Thanks to these samples, it was possible to compare theirforaminiferal contents with that from the Parnietowo boring, and so define more correctlythe age of the "tuffeau" series in the Pamietowo section . It was also possible to establishthe relation between the boreal Scandinavian Lower Paleocene (Selandian) sediments and themeridional 1 (non Mediterranean) west European Lower Paleocene (Montian) sediments fromtheir stratotypes.

THE BOREAL PROVINCE IN EUROPEAND THE REGIONS WITH SIMILAR BIOFACIES CONDITIONS

(Text-figs. 3, 4: Tables 1. 2)

Foraminifera occurring in the sediments of the Pamietowo boring are of Paleocene(Montian) age. From the point of view of ecology , the Foraminifera from the Pamietowoboring show that, in Northern Poland, the two different environments: boreal and meridional,merge. The boreal influence was somewhat stronger at the base of the Paleocene series (ascan be seen in the basal sediments of this boring) with the meridional (non Mediterranean)influence increasing later (top sediments of the Pamietowo boring). On the whole, however,the meridional influence is much weaker than the boreal. This conclusion can be drawn fromthe proportion of a little warmer to somewhat colder species, as well as from the state of theirtest development. In the Pamietowo boring boreal species predominate, subtropical species for­ming only 15%of the whole. Moreover the tests of the last species, in comparison with ana­logous species in optimal conditions, are much smaller, less variable, with weaker ornamen­tation, if at all, and thinner.

In the Tertiary sediments in the lower part of the Pamietowo section, there occurs anassemblage of Foraminifera similar to the well known assemblage, described from the LowerTertiary more sandy sediments of Central Poland (BROTZEN & POZARYSKA, 1957; POZARYSKA,1965). In this assemblage benthonic foraminifers predominate, especially representatives ofthe families: Cibicididae, Nonionidae, Polymorphinidae, Discorbidae and Anomalinidae.They, together with the sediments in which they appear, prove that the depth of this earlyTertiary basin was not considerable. The planktonic forms indicate clearly that this shallowsea was connected with the open sea. However, there are some clear differences between theforaminiferal assemblage in the Lower Tertiary sediments of Pamietowo and this describedfrom Central Poland, the main difference being the lack in Pamietowo of some rather largeForaminifera of the genera: Palmula and Planularia. The genus Robulus is very scarce in Pa­mietowo, just as are the index forms for Selandian such as Lamarckina rugulosa PLUM.and Ceratobulimina tuberculata BROTZ. A similar difference can be seen between the fora­miniferal boreal assemblage of Pamietowo and that characteristic for the Paleocene sedi­ments of Denmark and Sweden (Scania). On the whole, however, the majority of speciespresent in the Pamietowo boring, as well as in Central Poland, also occur not only in thePaleocene of Denmark and Sweden, but also in the Paleocene in Austria, England, Europeanand Asiatic regions of U.S.S.R., North and Central America (U.S .A.) and Australia. Those

1 The here used expression ..meridional" has the same meaning as in DEROO'S paper (1966).

APPROXIMATE EXTENSION of LOWERPALEOCENE SEDIMENTS in EUROPE

r::::~ Boreal Province ("Selondian Beds")[>: :« ~>l Meridiona I Province ("Montion Beds")r ~ directions of microfauna migration

o 100 200 ~OO 400 500km• ! ' I

4;

Fig. 3

pv- ---

-: ';-. -

16 KRYSTYNA POZAR YSKA & JANINA SZCZECHU RA

common for am iniferal elements are less nu merous in the Paleocene of Germany, S Hollandand NE Belgium (Limburg Province), and still less numerous in SW Belgium (Mons Basin)and France (Pari s Basin). These latter areas shou ld be considered as a different province - atr op ical or subtropica l one, i. e. meri diona l (non Mediterranean) one. The extent of thi s lattera rea can be seen by the geog raphical distribution of the warm-water assemblage of Foraminifera .

DENMARK AND SWEDEN

The Upper M aastrichtian (Stevnsian ) sediments in Denmark and Sweden are coveredby Danian sediments, developed as limestone, similarly as the below lying Cretaceous layers.Although th e Danian sediments were refer red by BROTZEN (1959) to Upper Cretaceous, in thepresent paper they are treated as the lowermost Tert iary sediments . In Denmark and Sweden,the beds younger than Dani an and old er than the lowermost Eocene, were designated byROSENKRANTZ (1924) as Selandian . These are det ritic, sandy-glauconitic sediments, lying onthe grey Upper Dan ian limestone. They begin as a basal co nglomerate, which lies unconformlyon the corroded hard groun d surface of Dan ian beds, or on flint layers, penetrating into thelatter's upper surface, being cut by numerous canals and burrows. This basal conglomerate isca lcareo us, conta ining numerous, small sized phosp ha tic nodules and small pebbles of Danianlimestone. The basal conglomerate of Sweden (Scania), the so-called Klagshamn conglo­merat e, is considera bly more calcareous than that of Denmar k, i. e. the Copenh agen conglo­merate. The latter contains a rich foraminiferal fauna, partly derived from the Danian, as wellas numerous badl y crus hed mac rofaunal remains, the majority of which are probably rework­ed . In Sweden (Scania), there is a thin series of green, calca reous, quartz-glauconitic sandwith numerous Foram inifera described by BROTZEN (1948), lying on an equally thin layer ofconglomera te (Kl agsham n qua rry near Malmo) .

In Denmark, Lower Paleocene (Selandian) sediments occur, in the first place, in Zealand(Lellinge, Copenhagen), as well as in severa l local it ies in Jutland (among others H valle se andEgsma rk). The microfauna of those sediments was described by FRANKE (1927) and latermentioned by BROTZEN (1948) when describin g Foraminifera from Sweden (Scania). Morerecently Selandian Foraminifera from Denm ark were quoted by BANG (fide L. B. RASMUSSEN,1960), and later supplemented by POZARYSKA (1965) an d HOFKER (1966). The below cited speciesbeing the mo st important, a re commo n to the Selandian and, in mo st ca ses, to the Danianbeds sediments of Denm ark, Sweden a nd Poland. Species not recognized in the Pamietowoboring, but occurring in Central Pol and, are marked by an as terisk. The list of species givenbelow seems to represent the most characteristic assemblage of foraminifers of boreal Paleocenein Europe. It should be menti on ed tha t ove r 40 % of those species listed occur al so in Denmarkin the marl known as .Kert eminde marl s" , described from Cop enhagen, the upper part of whichwas assigned to th e lower part of Upper Paleocene.

Spiroplectammina wilcoxensis CUSH. & PON. Globulina arenacea BROTZ.Arenobulimina cuskleya e JENN. Pyrulina fusiformis (ROEM).Robulus k lagshamnensis BROTZ.* Pseudopolymorphina frondea (CUSH.)Planularia pulavensis pulavensis POZ.* P. geijeri angusta BROTZ.P. discus (BROTZ.)* P. paleo cenica BROTZ.Astacolus gryi BROTZ. Sigmomorphina brotzeni HOF.A. paleocenicus BROTZ. S. soluta BROTZ.Palmula robusta BROTZ.* S. pseudoregularis CUSH. & THOM.

* Not known in the Pamietowo boring .

FORAMIN IFERA FROM TH E PALEOCENE OF POLAND 17

Nonionella ovata BROTZ.Elphidiella prima (TEN DAM)Bolivina oedumi BROTZ.Bulimin a ovata D'ORB.B. paleocenica BROTZ.Loxostomum applina e (PLUM.)*Pyramidina cuneata (BROTZ.)P. europaea (CUSH. & E DW.)

P. crassa BROTZ.Rosalina ko eneni BROTZ.R. ys tadiensis BROTZ.Latnarckina naheolensis CUS H. & T ODD

Gavelinella lellingensis BROTZ.Gyroidinoides pontoni BROTZ.G. octocamerata CUSH. & HANNAEponides toulniini BROTZ.

Osangularia cordieriana navarroana (CUSH.)Coleites reticulosus (PLUM.)Hocglundina scalaris (FRANKE)*Pulsiphonina prima (PLUM.)Ceratobulim ina tuberculata BROTZ.A labaniina niidwayensis BROTZ.Globigerina triloculinoides PLUM.Anomalina praeacuta VAS.A. ekblomi BROTZ.A. danica BROTZ.Cibicides proprius BROTZ.C. sahlstroetni BROTZ.C. simplex BROTZ.C. succedens BROTZ.Karreria fa llax RZEH.Tappanina selntensis (CUSH.)*

AUSTRIA

Austria , belonging to the Alpine geosyncline, is considered here because in the Paleocenebiofacies conditions were similar as in other countries of Boreal Province. The Foramini­fera , planktonic and benthonic, from the Paleocene deposits of Austria have been elaboratedin detail by HILLEBRANDT (1962) and also by SCHMlD (1962). Mention of some AustrianPaleocene Foraminifera was mad e by HOFK ER (1966) and POZARYSKA (1965). In a sampleof Paleocene sediments from Austria , from Krei sbach, north of Salzburg, kindly sent byProf. A. ROSENKRANTZ, the present writer s identified the following species : Robulus pseudo­mamilligerus (PLUM.), R. klagshamn ensis BROTZ., Angulogerina europaea CUSH. & EDw.,A. cuneata BROTZ., Pulsiphonina prima (PLUM.), Eponides toulmini BROTZ. , Bulimina midwayensisCUSH. & PARK., B. arkadelphiana CUSH. & PARK. , B. paleocenica BROTZ. , Vaginulina sp., Allo­morphina halli JENN. , Hoeglundina scalaris (FRANK E), Globigerina pseudobulloides PLUM., G. tri­loculinoides PLUM., Ceratobulimina tuberculata BROTZ., Anomalina praeacuta VAS., Osangularianavarroana cordieriana (CUSH.), Cibicides succedens BROTZ., C. proprius BROTZ. and Globo­rotalia angulata (WHITE). Among the above mentioned species, only Globorotalia angulata(WHITE) was never found either in the Polish or Dano-Swedish Paleocene.

The lowermost Tertiary sediments in 'Eastern Austria are developed in the form of sandand sandstone as well as lithothamnic limestone. The Foraminifera assemblage described bySCHMlD (1962) from the .Bruderndorfer Feinsand", referred by him to Danian, in reality seemsto represent Paleocene (Montian), older however, in all probability, than the Paleocene fromKreisbach, examined by the present authors. SCHMlD did not find the Paleocene (Montian)index form s, especially Ceratobulimina tuberculata BROTZ. , however , according to the presentwriters , species such as Robulus pseudo-mamilligerus (PLUM.), R. cf. klagshamnensis BROTZ.and Loxostomum applinae (PLUM.) confirm the Montian age of the "Bruderndorfer Feinsand".

ENGLAND

The oldest Tertiary sediments in England are of Thanetian age. Thanet Beds occurringin SE England, in Kent, are about 30 m thick and consist of marl , clay, silt, glauconitic-sandysilt and silty-sand, which in HAYNES' opinion correspond in age to the Lower Landenian ofPa l ieontolog ir, Palon ica No. ~ o 2

FRANCE I BELGIUM I HOllAND I GERMANY I POLAND I DENMARK I SWEDEN I U.S.S.R .• • • • • • • 1 ° "

meridiona I . . . . .-' ....:-:;: ::::.::<> :::.::

<I z foraminiferal

~,i 11et:o I .q~.~embla. _ .. . .

zCl')c:J ­o!:C.... uuCi;5t;w.J<(XcUz:

~•

...

lIIIIIIIll orqano-detritic limeslone, I :::':-::::~ quiucomiic calcareous sand, --.....~ direction of migration.

Fig. 4. Repartition of Foraminifera in the Paleocene of Europe.

f ORAMINIFERA FROM THE PALEOCENE OF POLAND 19

North Europe. Foraminiferal microfauna described by HAYNES (1954-1958) from the ThanetBeds, ju st as th at exa mined by the present a utho rs from th ese same layers (samples fromPegwell, horizons 47 and 54), shows conside rable similarity to tha t of th e Dani sh-Swedisha nd Polish Selan dian. This similarity confi rms th at there existed a connection and communi­ca tion between the Swedis h-Danish and Eng lish Paleocene basins .as well as between Belgiana nd French T hanetia n, as suggested by BROTZEN, 1948). The below mentioned species fromPegwell, commo n to the Eng lish, Polish a nd Dani sh-Swedish Paleocene, predominate in theassemb lage : Cibicides sahlstroemi BROTZ., C. proprius (BROTZ.), C. succedens BROTZ., C. cari­natu s (TERQ.), Anomalina praeacuta VAS., A. ekb/om i (BROTZ.), A. burlingtonensis (JENN.),A . umbilicata umbilicata BROTZ.. A. danica (BROTZ.), Anoma linoides nobilis BROTZ., Pyramidinacrassa BROTZ., Angulogerina europaea CUSH. & EDW., Osangularia cordieriana navarroana(C USH.), Globigerina triloculinoides PLUM. , Eponides toulmini BROTZ., Alabam ina midwayensisBROTZ., Rosa/ina selandiana n. sp ., R . ystadiensis BROTZ., R. k oeneni BROTZ., Bolivina oedumiBROTZ., Tappanina selmensis (CUSH.), Bulimina ovata D'O RB., B. pa/eocenica BROTZ., N oniongraniferum (TERQ.), Ceratobulimina tuberculata BROTZ.. Pulsiphonina prima (PLUM.), Episto ­minella cf. litnburgensis (VISS.), an d other s.

U.S.S.R. (EUROPEAN AND ASIATIC PART)

In the European pa rt of U.S.S.R. (Crimea excluded) the oldes t Tertiary sedimentsare of Paleocene age . Paleocene sediments occur in the depressions of the Ru ssian Platformin many regions, the most important of which are: Lithuania as well as Dni eper-Don etz Basinand ot her places of Ukraine. On th e Ru ssian Platform the Paleocene sediments are developedin th e so-called platformic facie s, correspo nding in cha rac ter to th e epicontinental faciesoccur ring in Northern Europe .

The Danian sediments occur in the south-east part of the Ru ssian Platform in theregion Ob shchyj Syrt, as well as eas tern of the U ra l River region. They represent a ratherthin seri es of limestones, 5- 8 m of thickness.

Marl y silts interbedded by marl s and sandston es, 15- 20 m of thickness, forming so­called "Tz iganovskij horizon " , are lying on th e Danian sediments in the Obshchyj Syrt region.This "Tz iganovskij horizon " is of Lower Paleocene age (LEONOV, 1961), i.e. Montian age ,or still of Dani an age (BEZRUKOV, 1933). Simil a r series are known also in th e Baskunchak­lak e region and in th e borin g at Novou zensko cit y.

In the southern pa rt of th e Russian Plat form, th e silty sandstones with glauconite,forming " Belogrodny hori zo n" of the Middle Volga R egion , and glauconitic sandstone of" Berezovs kij horizon" of the Lower Volga Region , a re of Lo wer Paleocene age (LEONOV,1961), i.e . Lower Montian.

" K anevskij hori zon" , developed as sandsto nes an d silts, is of Upper Montian orLo wer Tha netia n age (V. P. VASILENKO, 1950) and was found in borings at the DnieperBasin, between cities Rom ny and Zmijevym.

Of the U pper Paleocene age are: " Syzranskij horizon" , developed as " opoka"-rock,and sandstone of M iddle and Lower Volga Region, " Upper Tacinskij horizon" (Laoxov ,1961) - silty "opoka" -rock and silts of Don R iver Basin, and " Reshetn ikovskij horizon"(V. K . VASILENKO, 1946) - "opoka" -rock of th e Middle Desna region and of Upper NorthD on etz region.

Th e Paleocene sediments (Dania n and Montian) are developed in a similar platformic

2·

20 KR YSTYNA POZARYSKA & JANINA SZCZECHURA

facies in the Crimean Peninsula, Central Georgia and the Tadzhik depression (where Danianis lacking).

Lithuania. - The most northern region, in which the Paleocene sediments of the RussianPlatform occur, is SW Lithuania. The Paleocene sediments of Lithuania are developed ascalcareous sand and sandstone with glauconite, lying directly on Maastrichtian or Campaniansediments, with a maximum thickness of over 60 m. GRIGELIS (1960), describing the foramini­feral assemblage in the Paleocene sediments of Lithuania, compared it with the known assem­blage of the same age from the Ukraine, as well as with that from Poland and Sweden, statinga very considerable faunistic analogy and defining the age ot these sediments as Lower Paleocene(Selandian). The following species present in the Polish and Swedish Paleocene were also citedby GRIGELIS (1960) from SW Lithuania: Bulimina (?) paleocenica BROTZ., Cibicides (?) lunatus(BROTZ.), Ceratobulimina tuberculata BROTZ. , Alabamina wilcoxensis Toulm., Anomalina da­nica (BROTZ.), A. praeacuta VAS., Cibicides lectus VAS., C. proprius (BROTZ.), C. succedensBROTZ., Karreria fallax RZEH.

Dnieper-Donetz Basin. - The Paleocene sediments of the Dnieper-Donetz Basin are simil­arly developed as those in Lithuania and the below mentioned W Ukraine. Its foraminiferalfauna was described in detail by V. P. VASILENKO (1950), who correlated the two lower horizonsof the local Kaniew substage of the Dnieper-Donetz Basin with the Lower Paleocene of Swedenand Denmark. According to VASILENKO, these horizons ought to be referred to the Montian.The list of Foraminifera given by VASILENKO contains many species (cited below) commonto the Paleocene of the Dnieper-Donetz Basin, Sweden, Denmark, and Poland: Globulina gibbaO'ORB., G. amygdaloides REUSS, Reussella paleocenica (BROTZ), Alabamina wilcoxensis TOULM.,Gyroidina ex gr. soldanii O'ORB., Anomalina midwayensis (PLUM.), A . danica (BROTZ.), A. prae­acuta VAS., Cibicides(?) lunatus (BROTZ.), Cibicides lectus VAS.

Ukraine. - In the southern region of the Russian Platform, i. e. in W Ukraine, thePaleocene sediments do not exceed 70 m in thickness. KAPTARENKO-CERNOUSOVA et al. (1963)described from there an assemblage of Foraminifera, among which are the following species,known also from Poland and Sweden: Spiroplectammina variata VAS. cf. laevis (ROEM.),Gaudryina rugosa D'ORB. , Lenticulina (Astacolus) gryi BROTZ., Robulus discus BROTZ., Reussellapaleocenica (BROTZ.), Siphonina prima (PLUM.), Diofrondicularia geijeri (BROTZ.), Anomalina(Anomalina) danica BROTZ. , A . (Pseudovalvulinerla) praeacuta VAS. , Cibicides [Cibicidoides)lectus VAS., C. (Gemellides) lunatus (BROTZ.).

Crimea. - The lowennost Tertiary sediments lying on the Cretaceous in southern Crimea(Bakchisaray near Simferopol) are well developed and particularly well exposed, as an almost60 m thick deposit of the so-called Dano-Montian series. Most Soviet geologists and palaeon­tologists included it into Tertiary, only a few considering its lower part (Danian) as the Cre­taceous. It is a series of calcareous detritic limestone, sandy-marly at the base , passing upwardsinto organo-detritic limestone, similar to the detritic sediments of the same age in LimburgProvince, the so-called "tuffeau". The top of the series is developed as hard limestone withflints and siliceous marls, corresponding to calcareous gaizes. Mainly on the base ofechinids and foraminifers, this series was divided in the Crimea into Danian and Montian.The microfauna, characteristic of Dano-Montian sediments in the Crimea, was described byMOROZOVA (1959-1961), SUCKAJA (1958), MASLAKOVA (1959) and others. It is characterizedby a foraminiferal assemblage very close to that known from lowermost Tertiary, i.e. Danian

FORAMINIFERA FROM THE PALEOCENE OF POLAND 21

and Montian of th e Pol ish Lowlands. The lower part of th e discu ssed series represent s the Dan­ian, well characterized 'by index macrofauna, as well as by a n assemblage of smooth, planktonicForaminifera, while the upper represents Montian, its foraminiferal assemblage correspondingto the known asse mblage of Lower Paleocene (Seland ian) of Denmark, Sweden and Poland(MOROZOVA, 1960; POZARYSKA, 1965). MOROZOVA listed from the Montian of Crimea severa lspecies of Foraminifera, evidence of th e similarity between th e Selandian of above mentionedcountries. These are following the mo st important species : Ceratobulina tuberculata BROTZ.,Loxostomum applinae (PLUM.), Planularia discus (BRoTz .), Globigerina varianta SUBB. , Cibi­cides lectus VAS.

In samples from the Montian of Kursko e (Crimea ), th e present authors sta ted, in ad­dition, the presence of th e following for aminiferal species : Alabamina midwayensis BRoTz .,Eponides frankei BROTZ., Anomalina danica BROTZ. , A . praeacuta VAS., Osangularia cordie­riana navarroana CUSH., Bulimina ovata D'ORB., Cibicides succedens BROTZ. and Epistominellacf. limburgensis (VISS.) .

In the Montian assembl age of Foraminifera from Crimea, MOROZOVA (1960) also statedthe presence of the first represent at ives of Acarinina angulata (WHITE), not known from theSelandian bed s of North a nd Cent ra l E uro pe .

Georgia. - On the European-Asiatic transitional territories of U.S.S.R. , mainly inWestern Georgia there occur Upper Cretaceous and Lower Tertiary sediments, developed inplatformic facies. These are found on th e tectonic blo ck , situa ted between the Great and SmallCaucasus. The sed iments dep osited during the Upper Cretaceous and Lower Tertiary lie herealmost horizontally, not disturbed and never reaching a great thickness. The Danian sedimentslying conformably on the Maastrichtian , continue the same type of facie s, whereas the Pa­leocene sed iments lying non con formabl y with the Danian sediments, are eroded entirelyin several places. Often it lack s completely th e Lower Paleocene sed iments there.

The Upper Cretaceous and Lower Terti ary sed iments were examined there in the firstplace by TZAGARELI (1954) a nd KACARAVA & K ACARAVA (1960) . The Danian sediments ass ignedby Georgian authors to th e Cr etaceous, consist of white limestone, continuing th e Cretaceousca lcareous sed imenta tion. The higher lying deposits of green, glauconitic mar! and sandy siltcontain a foraminiferal assembl age close to th at in the Paleocene sed iments of Central Europe.However , they have numerous elements typical of geo syncline, for instance nummulites, whichhinder both the making of a close correlation between these distant sea basins and establishingits exact age, which probably should be placed not so low in strat igra phical schedule (POZA­RYSKA, 1965) as was stated by Georgian geologists and palaeontologists. The white limestone,assigned by th em to Upper Danian, should be assigned to the Montian on the base of thefollowing for aminiferal specie s : Lamarckina rugulosa PLUM., Robulus degolyeri (PLUM.), R. pseu­do-mamilligerus (P LUM.) , Hoeglundina scalaris (FRANKE) and Pyrulina fusiformis (ROEM.).

The abo ve cited species are never met with in European epicontinental Danian sediments.In addition, according to some Georgian author s, in th e layer considered by them as Danian,there occur also some Globorotalia s~~£!e;1Itft ~~~ as G. angulata (WHITE) a nd G. conicotruncata(SUBB .t~ in other region s datesthee,uWer" Paleocene bed s (BOLLI, 1966). Because of theoccurrence in the se layers of the above mentioned for aminiferal species, not known in depositsolder than Montian, and of others kn own only from Upper Paleocene in epicontmental as wellas in geo synclinal facie s, it seems that these layers should be assigned to the Montian and notto Danian, which in addition is considered in Georgia as belonging to the Cretaceous. Theforaminiferal assemblage reveals a great microfaunal similarity between the Paleocene of

22 KRYSTYNA POZARYSKA & JANINA SZCZECHURA

Georgia, Sweden, Denmark and Poland, showing an open connection between the epiconti­nental sea basins and tho se forming connecting basins to the open ocean s and great geosyn­clinal seas.

Tadzhik depression. - The Paleocene sediments of the Russian Platform are prolongedinto the Tadzhik depression, north of Amu-Daria , beyond the Caspian Sea. There the Paleoceneoccurs in the form of mar! and dolomite with gypsum. It was described by BYKOVA (1953)and later by Monozovx, KREJDENKOV and DAVIDZON (1965) and at the same time characterizedby Foraminifera. The Paleocene sediment s in thi s region were divided by the latter authorsinto three local horizons. The middle horizon, so-called "Aruktauskij Horizon" corresponds,in the opinion of these authors, to the European Selandian, however it was placed by theminto the Thanetian. The following species of Foraminifera , occurring there, are also commonto the Paleocene (Montian) of Poland: Lenticulina degolyeri (PLUM.), Gyroidina octocamerata(CUSH. & HANNA), Allomorphina aff. halli JENN. , Alabamina midwayensis BROTZ., Globigerinatriloculinoides PLUM. , G. pseudobulloides PLUM. , G. trivialis SUllD., Anomalina umbilicata (BROTZ.),A. ekblomi BROTZ., A. danica BROTZ. , Cibicides aff. succedens BROTZ. , C. burlingtonensis JENN. ,C. simplex BROTZ., C. proprius BROTZ., Bulimina ovata D'ORll., B. paleocenica BROTZ. andEponides aff. lunatus BROTZ.

Together with this assemblage, are found representatives of the Globorotalia angulata(WHITE) and G. pseudomenardii BOLLI , which are species, according to BOLL! (1966), character­izing the Upper Paleocene.

NORTH AM ERICA

In addition to the European continent, the boreal type of Paleocene sediments are knownfrom North and Central America and Australia. The foraminiferal microfauna occurring thereproves the existence of some connections in the Lower Tertiary between Europe, Gulf CoastalRegion of U.S.A. and Australia. In North America , the sediments assigned to the so-calledMidway Group correspond to the European Paleocene. These sediments were first characterizedin detail by H. J. PLUMMER (1926) and later by CUSHMAN(1944-1951). The Midway Formation.especially well developed in the Gulf Coastal Region , where the sediments reach a thicknessof almost 200 m, consists of clay, glauconitic sand, silt and limestone , all with concretions.At first, all the Midway Formation was compared with the Eocene (PLUMMER, 1926), later withthe Danish-Swedish Paleocene (F . B. PLUMMER, 1932) and lastly with European Danian (SCOTT,1934, fide BROTZEN, 1948, p. 32). In 1948, BROTZEN, on the base of Foraminifera, referred theLower Midway to Danian, and Upper Midway to the Dani sh-Swedish Lower Paleocene (Se­landian).

The present authors have examined a sample from the Upper Midway, Wills Point, MexiaClay member, from Texas. The below mentioned foraminiferal species present there, are alsoknown from the European Paleocene, this indicating the considerable similarity between theUpper Midway and Danish-Swedish and Poli sh Paleocene (Selandian): Robulus degolyeri(PLUM.), R. pseudo-mamilligerus (PLUM.), Pulsiphonina prima (PLUM.), Citharina plumoides(PLUM.), Hoeglundina sealaris BROTZ. , Loxostomum applinae (PLUM.), Alabamina midwayensisBROTZ., Anomalina midwayensis (PLUM.), A. ekblomi (BROTZ.), Lamarckina rugulosa PLUM.,L. naheolensis CUSH. & TODD, Osangularia cordieriana navarroana (CUSH.), Globigerina trilo­eulinoides PLUM., G. daubjergensis BRON. and G. pseudobulloides PLUM.

FORAMINIFERA BROM THE PALEOCENE OF POLAND 23

This list may be enlarged by species common to European Paleocene which have alsobeen described by CUSHMAN (1951) from American Paleocene. They are not cited here, becauseof their less stratigraphical value.

AUSTRALIA

Foraminiferal species common to both the North American and European Paleocene werealso identified by MCGOWRAN (1965, 1966) in Au stralian (We stern Victoria) Paleocene.Australian Paleocene, developed mainly in th e form of clay and sand, is represented by PebblePoint and Rivernook Member Formations, belonging to the Wangerip Group. On the ba seof the benthonic Foraminifera, MCGOWRAN correlated the Victoria Paleocene with the LowerPaleocene of North Europe and North America, however, on the base of the planktonic Fora­minifera known from the Tethys, he considered the Pebble Point Formation as Middle Pa­leocene, and the Rivernook Member as Upper Paleocene. In th e Au stralian Paleocene, amongthe species of Foraminifera described by M c GOWRAN, the following are common to theAustralian, American and European Paleocene: Citharina plumoides (PLUM.), Siphonina(Pulsiphonina) prima (PLUM.) , Robulus discus BROTZ. , R. pseudo-mamilligerus (PLUM.), Cibicidinamariae (RUPERT lONES) , Karreria fallax RZEH. , Lamarekina rugulosa PLUM., Hoeglundinasealaris BROTZ., Angulogerina europaea CUSH. & EDw ., Bolivinoides oedumi (BROTZ.), Tappa­nina selmensis (CUSH.) .

**

*

Some specie s characteri stic for the Paleocene of Boreal Province occur al so in the Paleoceneof the Meridional (non Mediterranean) Province in Western Europe, i.e. in Limburg Pro­vince , Mons and Paris Basin. The number of boreal species in the Meridional Provincedecreases westwards, while at the same time undergoing a gradual reduction in the higher partof the tropical or subtropical Paleocene. In th e Meridional Province the tests of boreal fora­minifers are smaller and less numerous than in the boreal. On th e ba se of an analysis ofsamples from the lowermost Tertiary of Limburg Province (Geulhem; Vroenhoven), thepresent authors considered the below mentioned species (only the most important st ra tigraphic­ally) as the most cosmopolitan, common to the lowermost Tertiary (Danian and Montian) ofthe Boreal Province and the eastern boundary of the Meridional Province.

Coleites reticulosus (PLUM.)Eponides toulmini BROTZ.Planulina limbata BROTZ.Frondicularia biformis MARSS.Elphidiella prima (TEN DAM)Textularia plummerae arkansasana CUSH.A/abamina midwayensis BROTZ.Robulus turbinatus (PLUM.)

Epistominella cf. limburgensis (VISS.)Cibicides carinatus (T ERQ.)C. sahlstroemi BROTZ.C. simplex BROTZ.Karreria fallax RZEH.Anomalina praeacuta VAS.A. ek blom i BROTZ.Pulsiphonina prima (PLUM.)

According to the writers' own observations of samples from the older Paleocene(Montian) of Limburg Province, Mons Basin and in Paris Basin (the foraminiferal assemblageof latter basin mentioned by BROTZEN, 1948), the number of boreal foraminiferal species is even

24 KRYSTYNA POZARYSK A & JANINA SZCZECHU RA

less numerous, while there is a rise in the number of meridional species which are moreoverbetter developed.

In Western Europe, only in th e Than et Beds the bor eal foraminiferal fauna a ppea rs asan almost exclusive assemblage.

THE MER1DIONAL (NON MEDITE RRANEAN) PROVINCE

(Text-figs . \ 4 ; Table 2)

As was ment ioned previously, in the lower part of th e Pami etowo boring an assemblageof boreal Foraminifera predominat es, however , single for am inifers chara cteris t ic of subtropicalPaleocene of West Europe also occur. In th e upper part of th e boring th e number of warm­water species increases, ho wever, they do not exceed 15 %of th e to ta l. The asse mblage of warm­water foraminifers is mainly represent ed by the fami ly Rotaliidae, being po or in other groupsof meridional foram inifers, especially rep resentatives of Miliolidae. The compositio n of th esubtropical foraminiferal assemblage from th e Pami ctowo bor ing, the test development of itsind ividual species, as well as th e relation in nu mber between the wa rm-wa ter and boreal species,all indicate that th e ecological conditions in th e Polish Paleocene were not sufficiently favourablefor the development of warm-wa ter for am iniferal microfauna . However , even thi s small per­centage of meridional foramin iferal fauna in the M ontian of North Pol and proves that in th isperiod the clim at e had becom e somewhat wa nne r in this part of Europe. On the other hand ,it sho ws th at at th at tim e th ere existed a co nnect ion between th e meridional sea of Westernand th e boreal sea of Eastern Europe .

The asse mblage of subtropical species occurri ng in the Parnietowo boring is known fromthe Lower Tertiary (Paleocene) of F rance (Pari s Basin), SW Belgium (M ons Basin) , th e Belgian ­Holland frontier (Limburg P rovince) an d th e weste rn part of Germ an y (K refeld region). Thisregion, in the upperm ost Cretaceous a nd lowerm ost Tertiary was occ upied by warm sea , inwhich the organo-detritic sed iments of "t uflcau" type predominat ed.

FRANCE (PA RIS BASIN)

The mo st classical area of th e Meridional (non Me dite rra nean) Province is th e Pari sBasin. The Paleocene (Monti an ) sediments a re developed there in th e form of limeston e,the so-ca lled "Calc aire pisolith ique" , which pro ba bly represents the o ldes t Tert iary sedimentsin thi s region. The age of "Calcaire pisoli th iqu e: was recog nized by M EUNI ER ( 1912) as D an­ian, by LEMOINE (1937) and ABRARD (1937) as Dan o-Montian , by MARIE (1937) and DAMOTTE(1964) as Montian and by van BELLEN (1946) as Danian or Mo ntian.

From the description of the fora min iferal asse mblage occurring in "Calcai re pisolithique" ,given by MARIE (1937-1964 and MS), it appea rs tha t the mo st predominatin g represent ati vespertain to th e follo wing families: Textul ariidae, Valvulinidae, At axophragmiidae, Mil iolid ae,Soritidae, Anomalinidae, Rotaliidae, D iscorbidae. Some of th e subtropical species men tionedby MARIE occur also in th e Polish Montian , the majority ho wever are common to Montianof Mons Basin and Limburg Province.

Th e present a utho rs had a t their disposal a single sample fr om "Calcai re pisolithiqu c'of Vigny , from which unfortunat ely only two meridi on al species, know n also from PolishMontian, could be identi fied . Th ese are Rota/hi trochidiformis (LAM.) and R . marginata D'ORB.

FORA MIN IF ERA F RO M TH E PAL EOCEN E OF POLAND

T A R LE 2

SOM E G UIDE FORA M IN IF ERA L ASSEM BLAGES IN T HE LOW ER PALEOCEN EW ANTAN AN D MO NTl AN ) O F EUROPE (T HE ALPIN E GEOS YNCLlNAL EXCLUD ED)

25

~~ ENGLAND fRANCE BElGIUM

cCl

HOllAt;o POLAND DENMARK SWEDEN U.S.S.R.

Epistomaric buruiensis Iv. Be/len)Rolorbinell a papilla ta n.soCuvittierina pomeraniana n.spNumerous species belonging 10

l1iliolidae famillj

Several new species f rom thePol Ijmorphinidae {ami IIjHol tandina peqwetlensis HaunesCib icides cti ssivel luun i HaunesCibituies cunobelini Haqnestibicides cantii Haljnes

tituc ides osteroiaes n.sp.6lobor% ha qtobiqerioitomus

[v. 8ellen jGlaboro/alia praepseudomerardii

HorAerR% rbine/la mariei (v. Bel/en)Rotorbinella montiona n. sp.vacuovatvutina keijzeri

Iv. Beueo)Ra/alia marqinata d'oiblgnlj

Species ex. gr. : Botivinouies, Stensioina ,Globo/r uncona. Rugog/abigerwa . Boliv in a,Neottabeu in a .

w ...Q)

ces:

Z 0-

w

u c

Cl.-

0 ...c0 0'

I: ..... c

....-' ,...-

Cl)

enet

.-cCl La. .-c00

cd

Cl) .-:) ...0 s:....... .-u ....< .......... '".... C10:: C1u L

Citnciti es cu rillieri Rou rii toisNonion graniferum I Ter quem]Pljru!lna Fusiformi s (Roemer )

, Rosalt'na setandiana n. so.tibiciaes succetiens BroteenProtelphid ium subiaere (t en Dam)Globiger ina korlowskii 8ro/zen & Poiar'l3kaGave/inella lel/ingensis I Bro/zen)Robulu s deqotqeri Plumm erLoxos/omum app/inae (PLummer)Pyramid/na europaea (Cushman r. Etiwaras]i amarckina ruqulosa Plummertomarckina oatieoiensis Cushman &. IoddCeratobulimioa tubercutata BrotzenCitucides lectus Va silenkoS/gmomorphina pseudoreqularis Cushmanl. Thoma sCibicides aurourae Rouvillois

Bouu o madrugaensis Iusbmt: n £ Berm. Stoborotolites qronuiatus n. sp.Baldia cubensis Cushman 8. Berm Elpbidiel ta pri ma ten Dam

I IEpistominella cf. iimburqensis (Visser )Eponities touimini BrotzenButirnino paleocenica BroireriAnomal/na umbilicate umbilicate (Bra/zen)Pseudopolqmorpbira paleacenica Brat zenGtobiqerma comoressa Plu mmerGtobiqerina pseueoouttoioes PlummerGlob/ger /na trit ocu tircides PlummerGlobiger ina daubie rqeasis Bronoimanr:

Ptanutaria outaveos i s Poi ar yskaPtcnala n a discus [Brotsen)

I

Pararotaua tuber ­cutitera (Reuss)Rotctia t rocbid i­iormis itamarck]Roititta saxorumd·Orblgnlj

andlarge torormniterc

. -e-v-__--:..I ---.' _---..:_-_---:...._-.............:---_...:....-_--tteridtona! Province Boreal Province

26 KR YSTYNA POZARYSKA & JANINA SZCZECHURA

SW BELGI UM (MONS BASIN)

Corresponding to the "Calcaire pisolithique" of Fr ance there is in the Mons Basin theso-called "Calcaire grossier de Mons" ("Calcaire de Mons"), limestone lying on a similar seriesof "Tuffeau de Cipl y". While "Calcaire de Mons" is known only from wells, the "Tuffeaude Ciply" occur s also in outcrops. The age of the latt er two series has been under discussionfor many years. Some authors, e.g. CORNET (I 922), VINCENT (1930) , WIENBERG RASMUSSEN(1962) and DEROO (1966) assigned "Tuffeau de Ciply" to the Danian , while the above lying"Calcaire de Mons" was referred to Montian . Oth ers, e.g. LERICHE (1937), DE HEINZELIN andGLIBERT (1957, in the Stratigraphic al Lexicon) and MARLIERE (1958) considered both the"Tuffeau de Ciply" and "Calcaire de Mons" as Montian. ROSENKRANTZ (1964) put the"TutTeau de Ciply" in the Middl e Danian , according to WIENBERG RASMUSSEN, and the"Calcaire Grossier de Mons" , which rests conformably on the tuffeau , in the Upper Danian .

Foraminiferal faun a of these sediments was described by HOFKER (1960-1966) a nd listedby POZARYSKA (1966). HOFKER, on the base of Foraminifera, assigned .T uffeau de Ciply" and"Calcaire de Mons" to Paleocene, comparing them to "Q" and "R" Zone of Limburg Province.On the other hand , POZARYSKA who had at her disposal samples from the boring in Mons,stated that the foraminiferal faun a occurring there does not permit one to define the strati­graphical boundary between the "Tuffeau de Ciply" and "Calcaire de Mons". FurthermorePOZARYSKA (see also BROTZEN & POZARYSKA, 1961) stated that the foraminiferal faun a fromthe Bunde Beds in Limburg Province, described by VAN BELLEN (1946), was very similar to thatoccurring in the Montian of Poland. The below mentioned species (from boring in Mons)are common to the lowermo st Tertiary beds of Mons Basin and the Monti an of North Poland.The determination of some of the below listed species given by POZARYSKA (1966) has beenhere revised,

Rotalia trochidiform is (LAM.)R. saxorum D'ORB.R. marginata D'ORB.Pararotalia tuberculifera (REUSS)Rotorbinella mariei (VAN BEL.)R. montiana n. sp.

The above given list conta ins thespecies are excluded.

Rotorbinella papillata n. sp.Vacuovalvulina k eijzeri (VAN BEL.)Globorotalia glob igeriniform is VAN BEL. ,G. praepseudomenardii HOF.Anomalina m inor n. sp.Cuvillierina ? pomeraniana n. sp.

meridional species only, the boreal, cosmopolita n

NE BELGI UM AND S HOLLAND (LlMBURG PROVI NCE)

The lowermo st Tertiary of the Limburg Province is developed as a glauconitic tuff chalkthe so-called "Tuffeau glauconieux" passing higher into tuff chalk. On the base of the Ostracoda,DEROO (1966) correlated "Tuffeau glauconieux" with "Tuffeau de Ciply" whereas the tuffchalk, resting on "Tuffeau glauconieux" was referred. to "Serie interrnediaire" and "Calcairede Mons" of the Mons Basin. According to DEROO, "Tuffeau glauconieux" belongs to Danian ,and the overlying tuff chalk beds - to Montian. HOFKER (1966) divided the upp ermost Cre­taceous and the lowermost Tert iary layers into zones. To the uppermost Cretaceous, i. e. Dano­Maastrichtian, he included among others "N" (" Me") Zon e, i .e. "Tuffeau glauconieux", whilethe abo ve lying tuff chalk was assigned to Paleocene. Within the Paleocene of Limburg, HOFKERdetermin ed "P" Zone , correl atin g it with the Swedish Lower Paleocene (Selandi an), "Q"Zone , with "Tuffeau de Cipl y" and "R" Zone, with "Calcaire de Mons". In thi s way he statedthe analogy between the Lower Tertiary of Mons Basin and Limburg Province.

FORAMINIFERA FROM THE PALEOCENE OF POLAND 27

The above given divi sion of HOFKER of uppermost Cretaceous and lowermost Tertiaryin Limburg Province is based on foraminiferal fauna, which is described in detail by this author.VAN BELLEN (1946) , however, was the first to describe the tropical Foraminifera from the upperpart of tuff chalk from so-called Bunde Beds. He referred the Bunde Beds (HOFKER'S "R" Zone)to Middle Eocene. In 1947, however, TEN DAM stated, as earlier suggested by MARIE (1945),that there is a similarity between the foraminiferal fauna of the French, Belgian and DutchMontian .

The present authors, who had at their disposal numerous samples from the uppermostCretaceous and lowermost Tertiary of Limburg Province (Vroenhoven, Geulhem, Bunde),found there numerous boreal and subtropical foraminiferal species; among them all the subtropi­cal species present in the Montian of the Pamietowo boring and in the samples from the MonsBasin (see list on 'p . 26). An analysis of the vertical distribution of foraminifers in the lowermostTerti ary of Limburg, undertaken by the pre sent authors, showed that the number of subtropicalspecies and represent ati ves increase towards the top of the sections. A similar situation isfound in North Poland (Pamietowo boring), however in Limburg the warm sea tran sgressedalready in Upper Maastrichtian which accounts for the single specimens of warm-water Fora­minifera found there, e. g. Rotalia trochidiformis LAM. , R. saxorum D'ORB. and Pararotaliatuberculifera (REUss). The influence of meridional sea was not felt in North Poland before theMontian, therefore the warm-water foraminiferal species are lacking in older layers.

GERMANY

The lowermost Tertiary of Germany is, so far , little known. The Paleocene of NW Ger­man y was described in very general terms by STAESCHE and HILTERMANN (1940) and dividedby them into Lower and Upper Paleocene. The determination of the not numerous Foramini­fera, listed and partly figured by the se authors, is rather provisional. Judging from the notvery clear illustra tions, there are , besides the boreal type Foraminifera assemblage, a fewwarm-water foraminiferal species. A more uniform foraminiferal Paleocene assemblage fromNW Germ any (Krefeld region) ha s been described by INDANS (1965). Among the species givenby TNDANS, the follo wing are in common with the Paleocene of NW Germany, lowermostTert iary of the Meridional Province and North Poland: Te xtularia plummerae tLALICKER),Anomalina grosserugosa (GOMBEL) (recte A . danica BRoTz.) , Eponides gratus (Rsuss) (recteE. toulmini BRoTz. ), Discorbis corrugata (CUSH. & BERM.) (recte Rotorbinella montiana n. sp.),Rotalia saxorum D'ORB. , Coleites reticulosus (PLUM.), Rotalia trochidiformis (LAM.), Marsso­nella k eijzeri VAN BEL. [trecte Vacuovalvulina k eijzeri (VAN BEL.)], Elphidiella prima (TEN DAM).

In alm ost all the Meridional Pr ovince , the Montian beds are covered by lagoonar sedi­ment s, conta ining a boreal type of Foraminifera and are usually correlated to English Thanetian.Th e foraminifers of the se sediments, similar to that of Danish-Swedish Paleocene (Selandian) ,have been presented in a pre vious part of thi s paper.

THE STRATIGRAPHY OF THE LOWERMOST TERTIARYIN EUROPE

(Table 3)

In the previou s part of thi s pap er , when the lowermost Tertiary in Europe was being ana­lysed , attention was concentra ted mainly on the differences in the ecological conditions inEurope in Lower Terti ary and , connected with thi s, the differentiation of foraminiferal micro-

28 KR YSTYNA POZARYSKA & JANI NA SZCZECHURA

faun a, while the stra tigraphy of thi s period in Euro pe wa s tre ated onl y in general terms. Thi swa s because the problem of the st ra t igra phy of lowermost Terti ary, just as that of the Cretaceous­Tertiary boundar y, has not , up to now, been definit ely resol ved . Therefore particular, loca lhorizons were ass igned to different strat igraph ical un its. Especially co ntroversia l is the stra t i­graphical po sition of the Danian and the correlati on of local Tertiary horizons in the area inquestion .

The Danian stage was established by D ESOR ( 1846) who considered th e limestone fromFakse in Zealand as typ ical of th e Dan ian , com paring it to th e lime stone from Laversine inFrance, i. e. the . Calcaire pisolithique" of th e Pari s Basin. However, the results of further in­vestigations on the "Ca lcai re pisolithique" of France showed it to be of the same age as "Calcairede Mons" from Belgium, i.e. th e st ra to type of Montian (CORNET & BRIART, 1865) . As a result,the typical sect ion of Dani an was regionally restricted to the territory of Denmark, being lo­cali sed th ere in Fak se in an old quarry on land, where, as it later appeared, only Middle Danianlimestone occurred. Borings in Faxe sho w the Lower Danian on top of the Maastrichtianas well. Th e typical sect ion was completed by sho re expos ures at Stevns Klint representingLower Dani an , and further extended by outcropping in th e neighbouring region of Sweden(Scania), where, in add it ion to Lower a nd Middle D ani an , the Upper Danian is also repre­sented . Upper Dani an is pre sent also in East Zealand, Copenhagen , Funen and Yutland .TROELSEN (1956) a nd BROTZEN (1956) (fide BROTZEN, 1959) designated the above mentionedexpos ures of East Zealand, together with th ose in Scania, as a typ e-r egion , representinga complete D an ian profile from the lowermost to uppermost inclusive.

In contrast to the ori gina l stratigraphical position of the D ani an, defined as uppermostCretaceous byDrson (1846), DE GROSSOUVRE (1897) assigned it to the Tertiary as its lowermosthorizon. However , it appeared th at much ea rlie r than either DE GROSSOUVRE or DESOR, theCerithium lime ston e of Stevns Klint an d Bryozoan limeston e with Coral lime stone from Faksewere described by FORCHHAMMER (1825), on the base of the molluscs fauna compared with"Ca lca ire pisolithique" from the Pari s Basin , and ass igned by him to the Tertiary. Next toaccept Ter tiary age for th e Dani an was BRONNICH NIELSEN (1920) , who moreover includedDanian into th e Paleocene, ROSENKRANTZ (1920) a nd HARDER (1922) ; th is view being heldup to the well kn own paper by RAVN (1925) , the latter placing it ba ck again into the Cretaceous.

U p to no w, the stra t igra ph ica l po sition of th e Danian is controver sial. In spite of manyworks o n th is subjec t, the Dani an is sti ll con sidered by some authors as belonging to theCretaceous, while others assign it to th e Tertiar y. At present , in Denmark the prevailing opinioninclines to wards the Tertiary. On the other hand , in the neighbouring territory of Sweden, thetradit ion al view is still held , i. e. th e Danian is assigned to Cretaceou s (BROTZEN, 1959; see alsoLex. Strat igr. Intern ., I , Europe, 2, p . 87, M exico). BROTZEN mentioned certain differencesbetween the Upper Dani a n mic rofa unal assemblage and th at from th e Middle Danian. Henoted that th e newly appea ring microfauna of Upper Danian had already the character ofTertiary. An ana logous view was lat ely given by BANG (per s. comm.) about th e appearance ofPaleoc ene for am iniferal asse mblages (Seland ian ) in th e uppermost Danian sed iments in Den­mark. The problem of whether th e Danian sho uld be assign ed to the Cretaceous or Tertiary,was also recently d iscussed by BERGGREN ( 1964).

Up to now , there is no mon ographical elabora tion of the Foraminifera either from thestra totype of th e D ani an in Denmark or from the Danian in Sweden (Scania), i.e. from thetype-regi on , with th e exception of BROTZEN'S remarks (1948, 1959), TROELSEN'S (1957) andBANG'S remarks (1960 , f ide RASMUSSEN, a nd 1962). A more complete list of Foraminifera fromthe Danian of Denmark, with illu strations and remarks, was given by HOFKER (1966). The

Palaeontologia Polonica, No . 20, 1968 K. Poi aryska & J. Szczechura, 10/ . 3

T AB LE 3

DI AGRAMMATIC CORRELATION O F THE MOST IMPORTANT SECTIONS OF THE LOWERMOST TE RTIARY lN MIDDLE EUROPE

ENGLAND(Kent)

F RAN CE(Par is Basin)

BELGIUM(Mons Basin)

BELGIUM-HOLLAND(Limburg)

POLAND(Carpa thians excluded)

D ENMA R K(Zealand)

SWEDEN(Scania)

U. S.S. R.(Russian Platform)

~ - ---- -- ------ --

I .1- -- - 'II UJ

(J

<t:

(lI AY NES. 1954) (MARIE, 1937-64.ROUVILLOlS. 1960.

DAMoTTE, 1964)

(MARLIERE, 1957-62 ,HOFKER,1966 ,

POZARYSKA, 1966)

(VAN BELLEN, 1.946,MEtJER 1959,DEROO, 1966,

HOFKER, 1966)

(BROTZEN & POZARYSKA, 1961,POZARYSKA, 1965,

POZARYSKA & SZCZECHURA,present paper)

(ROSENKRANTZ, 1924-60,BROTZEN, 1948,

BANG, 1960)

(BROTZEN, 1948, 1959) (GRIGELIS, 1960, K APTA­

RENKO-CERNOUSOVA, 1963, iVASILENKO, 1950, I

Monozovx, 1965-66, ILEONov, 1961)

--_.- - - - - -1- -- --- - - - - - - ------- - - - - -

Wool wich and Reading .

Beds

T hanet Beds(Th anet s. s.)

*)

Thanet Bedssand limestone

Freshwater sand with ligni te, HOFKER'S S ZoneHeersian limestone and sa nd

Missing

" Kerteminde mar l"

I

In erra tic blocks k~wn as"Maglehem bou lders"

I

Upper Tacinskij horizon;siliceous marl, silt, and

Syzr anskij horizo n ; opokaa nd sa ndstone

Bryozoan limestoneI

Co ral limestone

Bryozoan lime stone

Danian s.s, = Cora l andbryozoan lime stone

Kanevsk ij horizon; sandyglauconitic silt

Belogro dny and Berezo vskijhorizons ; marl y sandsto ne,

SeJandian s.s . = Lellinge green- Selandian s.s. = Klagshamn sand O bshchij syr t ; Tz iganovskijPulawy Bed s sand (Pa leocene of Copenhagen) and clay ! horizon; silt , marl , I

I sa nds to ne 1'so-ca lled "siwak" ! I

_ _ _ I _,~~o,s~~~:~t i~ _ ~on~Iome~t~__~~::~glomerate~~_ I limestone I(t hin series)

I Coccol ith ic limestone

Sochaczew Bedsmarly glauconitic san d

Pamietowo Bedsdetritic sandy limestone

" tuffea u" type

Bund e BedsHOFKER'S R Zone

TufT chalk

Tu fTeau glaucon itic = HOFKER'S N("Me") Zone

?

Ciply limestone"Tuffeau de Ciply"

Lacustrine and lagoonar Montian I= HOFKER'S S Zone i

Mons limestone I,.Ca lca ire de Mons"

(Mo ntian s.s.)Compact limestone

?

M issing

..de Vigny" reeflimestone

"Ca lca ire pisol ithique"

UJ

ZZ-c

LJJ 1=z~ 0

~

0I

LJJ i....I '-I<

I~

iII Z

I<z<Cl

- - - - -'-------- - --- - -- - - - - - - - -- - ---- - -'--

Missing

- , ~.- - -, ' -. ' ~' --" - - '- - --" - ' .- ' - - 1- - - - - - -

ChalkWh ite chalk (Stevnsian)

Fi sh clay

Cerithium limestone

White chalk (Stevnsian )Zyrz yn Beds

Siliceous mar! so-cal1ed "opoka"

TufTeau not glauconitic "Md" =

HOFKER'S L-M Zones.T uffeau SI. Symph oricn"

Co nglomerate" Poudingue de la Malogne"

.,

I

I

1- - -----Cha lk

! 1­i -~-- ~

UJ9 I: ~ ~UJ UU a.,-< a..~f- :::If-UJ I ~

~ I <U I ~

*) ~_ • . unconformi ty or h iatus.

fORAMINIFERA FROM THE PALEOCENE OF POLAND 29

latter author referred the Danish Danian to Cretaceous, including it to the Dano-Maastrichtian.Earlier HOFKER (1960, p. 53) based the correlation of the Danian of Denmark with the upper­most Cretaceous of Limburg Province on the evolution of pores in tests of several speciesfrom the Maastrichtian and Danian. This theory, however, was not supported by any micro­palaeontologist working on this problem .

In the present writers' opinion, the appearance in the Danian of Denmark and Swedenof foraminiferal assemblages characteristic for the early Tertiary (forarniniferal species citedby BROTZEN (1959, p. 23) and HOFKER (1966, pp . 294-304)), proves th at the Scandinavian Danianalready belongs to the Tert iary. The not numerous Cretaceous species pre sent in the Daniansediments of Scandinavia could be considered , as in Poland, as being reworked, or as disap­pearing Cretaceous relicts in the lowermost Terti ary. This last phenomenon may result from thefact that in the type-region, the Danian deposits are developed in facies somewhat similar to theunderlying Cretaceous. Po ssibly the conditions in the Danian sea were identical or very closeto those existing during the late Cretaceous, thanks to which the Cretaceous fauna had a betterchance of continuing into the Danian in thi s regio n, than in ot her, more eastern regions.At the end of Maastrichtian, true belemnites tBelemnella casimirovensis, in the area underdiscussion), ammonites (Hoploscaphites constrictus vulgaris, ibidem) and inoceramids (lnocera­mus tegulatus , ibidem) became extinct, in spite of the fact that the ecological conditionsprobably remained stable. The lack of micro-index forms in the Danian makes it difficultto correlate on a large area. The stratigraphy of the Danian bed s of Denmark and Sweden(Scania), based on Tylocidaris remains , could not be applied in other regions, because of thenearly total lack of the se forms of echinids and the type of facies with which thi s particularfauna is connected.

Danian bed s in Denmark and Sweden are covered by so-called Selandian beds. The Se­landian stratotype is described by ROSENKRANTZ (1924) from Lellinge in Zealand, south ofCopenhagen. It represents stro ngly glauconitic, weakly cemented calcareous sand, lying directlyon white Danian limestone. ROSENKRANTZ (1924-1960) assigned it, together with Danian, tothe Paleocene, i. e. within the frame of Tertiary. In the light of recent investigations, thi shas been fully confirmed. BROTZEN (1948), describing foraminiferal microfauna from the Se­landian of Sweden (Scania), referred it also to the Lower Paleocene.

The most recent studies in Poland (POZARYSKI & POZARYSKA, 1960; POZARYSKA, 1965),show that the Poli sh Danian in the Lowlands is confined to a very restricted area and. .£9CJlfSmainl y as remnants. All tho se places where the Poli sh Danian and Montian do not vary/if~y,are grouped along a line running from NW to SE Poland. Thi s line is an axis of the marginalsynclinorium. These sediments are preserved mainly in the Middle Vistul a Valley, in the vicinityof Pulawy (Bochotnica and Nasilow). . k. <:~e

Danian sediments in Poland are very strongly differentiated 'f~~~· In Poland, as on thewhole eastern European Platform, terrigenic type of sediments predominate, such as: sand,sandstone, marl, calcareous gaize with glauconite and so on. The Danian sediments in thePolish Lowlands are usually not of great thickness, from a few to about twenty metres. In manyplaces they are completely eroded, occurring only as remnants. Greater thickness is foundonly in eastern Pomerania, i. e. the north province of Poland.

The foraminiferal assemblage in the D anian sediments of Poland differs from that occurringin the underlying beds of Maastrichtian age (POZARYSKA, 1952, 1965). In addition to the notnumerous Cretaceous for aminiferal species occurring in the Danian, where they are ratherreworked, the majority of species appearing in the Danian are continued into the above lyinglayers of Montian age. Thi s is why the Danian in Poland is con sidered as definitely of Tertiary

30 KRYSTYNA .POZAR YSKA & JANI NA SZCZECH URA

age (POZARYSKA, 1965) and placed into Paleocene (POZARYSKA, 1966). Because of the absenceof index forms characteristic exclusively for Danian, assignment of the Danian to Paleoceneis based on a negative premise, i. e. on the lack of certain elements characteristic for the higherlying deposits of Montian age. The ostracods examined by SZCZECHURA (1965) did not permitdifferentiating the Danian from the Montian so as to establish the Danian as a separate strati­graphical unit. They support, however, the position of the Cretaceous-Tertiary boundary, as lyingbetween Maastrichtian and Danian, which was previously stated on the base of many animalgroups, and finally on the base of Foraminifera.

Several Maastrichtian macrofaunal elements stated in Poland at the base of the Daniansediments or occasionally in Montian in places where the Danian is completely eroded, areundoubtedly reworked. This supports the supposition that Cretaceous microfauna in lowermostTertiary deposits can also be reworked.

In Poland, all the foraminiferal assemblage passes from the Danian into the Montiansediments, which as suggested by the present authors, correspond in age to those of the Selan­dian in Denmark and Sweden. The development of Montian sediments in Poland is analogousto 'that of the Selandian sediments from Denmark and Scania, all containing analogousassemblages of foraminiferal microfauna.

A correlation of lowermost Tertiary sediments of Denmark and Sweden, i. e. Danianand Selandian sediments, with those of Poland does not present any special difficulties. Inthe U.S.S.R., where the Danian is included by several authors to Tertiary (among othersKELLER, 1956; BYKOVA, 1956; NAJDIN, 1960; POSLAVSKAJA& MOSKViN, 1960; CHALILOV, 1948;MOROZOVA, 1960), the Danian sediments, especially those in the European part of U. S. S. R.,differ from the Cretaceous ones as well as from the higher lying Paleocene sedimentsknown as Selandian or sometimes as Thanetian. The lowermost Tertiary sections of the Euro­pean part of U. S. S. R. are easy, however, to compare with analogous series from Poland andScandinavia.

Whereas, there is no great difficulty in correlating lowermost Tertiary beds of Polandwith those of North and Eastern Europe, some difficulties arise when comparing the uppermostCretaceous and lowermost Tertiary beds of Poland with those of Western Europe. This resultsmainly from the fact that the facies of sediments of this age in Western Europe are very varying.Both in the Limburg Province and the Mons Basin , the Lower Tertiary sediments (Danian andMontian) are developed in the form of pure zoogenic limestone of the organo-detritic seriescalled "tuffeau". Foraminiferal microfauna found in these calcareous sediments is decidedlymeridional, in contrast to that contained in sediments of the same age in Denmark, Swedenand U. S. S. R., where it is definitely boreal.

The presence of the Danian in the Mons region and in Limburg is difficult to proveon the base of Foraminifera. Many authors, especially in Belgium and Holland, have concernedthemselves with this problem. Recently the problem was investigated in more detail by MAR­LIERE (1957, 1958, 1962), WIENBERG RASMUSSEN (1962), HOFKER (1937-1966), MARIE (1964),DEROO (1966) and others.

The Danian in the Mons Basin, i.e. the "Tuffeau de Ciply" series, was assigned to Danianby VINCENT (1930) and later by WIENBERG RAsMUSSEN (1962) and DEROO (1966). On the baseof MARLIERE'S supposition in an unpublished supplementing plate to his communication (1962),the "Tuffeau de Ciply" series was referred by him to Danian. However, in the text itself herather implies that the age of the .Tuffeau de Ciply" is the same as the above lying "Calcairede Mons", the latter being the stratotype of Montian. ROSENKRANTZ (1964) considered the"Calcaire Grossier de Mons" as Upper Danian. MARIE (1964) and HOFKER (1966) assigned

FORAMINIFERA FROM THE PALEOCENE OF POLAND 31

the "Tuffeau de Ciply" series to Montian , and not to Danian. All the above mentioned authorsexcept WIENBERG RASMUSSEN, are inclined to consider the .T uffeau de Ciply" as representingTertiary, as was suggested much earlier by RUTOT (1885) and VAN BROECK (1885).

The post-Maastrichtian sediments of Limburg Province are covered by a thin series,the so-called "Tuffeau glauconieux" ("Me" local horizon), occurring on a hard ground layerof Upper Maastrichtian age ("Md" local hori zon) ; they are present in Belgium in the AlbertCanal (Vroenhoven) as well as in neighbouring Holland (Geulhem). According to MEIJER(1959), WIENBERG RAsMUSSEN (1962) and D EROO (1966), "Tuffeau glauconieux" is of Danianage and, in the opinion of the last two authors, corresponds to the .Tuffeau de Ciply" seriesfrom the Mons Basin, as well as to the Danian from Denmark. Recently HOFKER (1966), whoalso compared this series to the Dani sh Danian , assigned this local " Me" horizon to the so-calledDano-Maastrichtian series, within Cretaceous. According to this author, only "P" zone shouldbe considered as the beginning of the Tertiary period.

In the present writers' opinion, the Cretaceou s-Tertiary boundary in the South Limburgarea should be placed between HOFKER's local " M" (upper part of " Md" horizon) and "N"("Me" horizon) zones, on the base of the distribution of foraminiferal species presented byHOFKER (1966, tabl e 1). Other authors, e.g. MEIJER (1959), WIENBERG RASMUSSEN (1962) andDEROO (1966), are of the same opinion as the present authors.

By comparing the microfauna of Denmark and Sweden (Scania) with that of WesternEurope on the one hand, and Eastern Europe on the other, the present writers are unableto distinguish the Danian in Western Europe. The foraminiferal microfauna characteristicfor the Montian sediments ("Calcaire de Mons") occurs also in the below lying "Tuffeau deCiply" series, as is shown by some authors (among them POZARYSKA, 1966). A few Montianspecies appear as early as in the Upper Maa strichtian ("Ma" hori zon) in Limburg Province.In spite of this the "Tuffeau de Ciply" series in the Mons Basin and the corresponding "Me"horizon in Limburg Province are only tent atively assigned to the Danian by the present writers,supporting the view of WIENBERG RASMUSSEN and D EROO.

It is proposed in this paper, that the position of the Danian in the general stratigraphicalschedule should, together with the Montian, be considered as Paleocene within the frame oflowermost Tertiary. In the included stratigraphical table, theDanian is shown under the Montian,to which are referred deposits more or less of the same age as Selandian. Because the Daniandiffers greatly in its foraminiferal assemblage from the Ma astrichtian, while varying onlyslightly from the upper lying Montian , and because in Western Europe (the Carpathians andAlps excluded) the Danian presence may be problematic, it is a question if the Danian doe smerit to be ranged as a sepa ra te stage.

CONCLUSIONS(Text-figs. 3, 4 ; Tables 2, 3)

From the investigations of the lowermost Tertiary Foraminifera, which have been carriedout up to now in Poland (the Carp athian s excluded), and their comparison with assemblagesof Foraminifera of the same age from other countries, the following palaeogeographic pictureappears.

In the Lower Paleocene of Europe, including Denmark, Sweden (Scan ia), Poland andthe European part of U.S .S.R., there was an extensive boreal sea basin strongly under the

32 KR YSTYNA POZAR YSK A & JA NI NA SZCZ ECH URA

influence of the North Atlantic Basin . This boreal basin did not represent a very extensive sea,but was composed of several elongated, narrow, intercontinental basins, having the characterof narrow straits between the surrounding continents, which supplied the sea with rich terri­genic material. Thi s elongated boreal basin cons isted of several individual sea bas ins, the mainbeing the Donetz Basin, Danish-Swedi sh Basin, Polish Basin, Crimean Basin and LowerVolgian depression . The character of its sedimentation varied according to the changes in theline of the sea shore. The Laramic synorogeny from the Cretaceous-Tertiary boundary stronglyinfluenced the character of this boreal, northern province as applied to its size and to the cha­racter of its sediments.

In this Boreal Province the strongest influence of the Laramic synorogeny was felt in thearea of Poland, where it gave rise to the upheaving of the Pomeranian-Kuiavian anticlinoriumin the lowermost Tertiary, i.e. in the Dan ian , which result ed in an inevitable regre ssion ofa shrinking Upper Cretaceous sea from the territory of Poland. On the other hand, on theDani sh-Swedish territory, compensatory movement s cau sed, in Danian, a subsidence of thebottom of this sea basin, a con sequence of this being an increa se of the sedimentation, as wellas a continuence of the zoogenic Cretaceous sedimentation, which existed up to then .

An analogical situation existed in the southern area of the Russian Platform , especiallyIII the part adjoining the Crimean peninsula , where in the lowermost Tertiary, i.e. .Dano­Montian", a calcaro-zoogenic sedimenta tion, very similar in character to Cretaceous, was con­tinued just as in Denmark and Sweden during Danian.

The foraminiferal microfauna, cha racteristic for Lower Tertiary sediments of the BorealProvince, is of benthonic character, with very meagre participation of planktonic forms (gro­bigerines only) and without meridional elements. Only on the periphery of the Boreal Province,especially the transition zone in the southern basins with geosyncline, among others, in theCrimea, meridional elements appear (SUCKAJ A, 1958), penetrating from the neighbouringgeosynclinal sea. On the other hand, a meridional province existed in the area of WesternEurope in lowermost Terti ary. It was probably connected at that time with the AtlanticOcean and acro ss France with the Tethy s Ocean. Thi s province, which can be called the Merid­ional (non Mediterranean) Province, occupied a comparatively small area of Europe, muchsmaller than the Boreal Province. It extended over the eastern part of the Paris Basin andMon s Basin, which at that time was a small bay of the form er, further east being narrowlyjoined to the Limburg Basin of Belgium-Holland and con tinuing somewhat eastwards to theterritory of the Lower Rhine in Western Germany.

The predominating type of sediments in the Meridional (non Mediterranean) Provincewas calcaro-detritic of the type "tuffeau" zoogenic sediments, similar to the reef sedimentsof a shallow, warmer sea. This type of zoogenic-detritic limestone was being con stantly de­posited in this basin right from the beginning of Upper Maastrichtian up to Montian. TheLaramic synorogenic movem ents in thi s area were much weaker, having only an insignificantinfluence on the type of sedimenta tion in compari son with its influence in Central and Ea­stern Europe.

Foraminiferal fauna occurring in the lowermost Tertiary of the Meridional (non Me­diterranean) Province, consi sts mostly of benthonic form s, being poor in planktonic forms,ju st as in the Boreal Province, however, here meridional elements predominate (see Table 2).The presence of boreal form s in the Dano-Montian foraminiferal assemblage of the Meri­dional (non Mediterranean) Province, as well as the presence of subtropical foraminifers inthe marginal western region of the Boreal Province in somewha t younger Montian is evidenceof the existence, in those times, of some sea conn ection between these provinces, which made

FORAMINIFERA FROM THE PALEOCENE OF POLAND 33

the exchange of microfauna po ssible. As Lower Tertiary sediments have so far not been foundbetween the above mentioned provinces, it is difficult to define exactly the route of the con­nection, however, an analysis of the regional and stratigraphical distr ibution of Foraminiferagives some idea as to th e general route and sequence of faunistic migration.

Microfaunistic int erchange, which too k place in the lowermost Tertiary between Eastand West, of Europe, p roves that the ecological conditions of the Boreal Province were unfa­vourable then for meridional fauna. Only in the younger Montian was the climate more favour­able, the result of the warming of the clima te of the western periphery of the boreal basin.This permitted a two-directional int erchange of fauna, i.e. from West to Ea st, as well as fromEast to West. POZARYSKI and POZARYSKA as early as in 1959 indicated that warm-water Tertiaryforaminiferal microfauna appeared earlier in Western Europe than in Eastern Europe, dueto the more favou rable ecologica l conditions. This applied, however, to somewhat older micro­fauna , i.e. Terti ar y forms appea ring in the Belgian-Dutch Limburg already in the UpperMaa str ichtian. Present investigati ons show that the earli er appearance of warm-water micro­fauna in Western Europe was not confined to Upper Cretace ou s, but continued also in thelowermost Terti ar y.

Analysing the Pa mietowo section and correlating it with other sections from CentralPoland , it is necessary to state that the main mig rati on of subtropical fo raminifers in NorthPoland took place in the yo unger Mont ian, when older Montian sediments of the Selandiantype, especially in Central Polan d, were already deposited. On the base of pre sent data it seemsthat the migration from the West to Eas t took place through German y, from where subtropicalMontian foraminifers were mentioned (INDANS, 1965) and that Poland represent s the mosteastern area under meri diona l (although very weak) influence.

In the opinion of the present au thors , the Dano-Montian sediments of the Meridional(non Mediterranean) Province should be laterally correlated to the Dano-Selandian sedimentsof Boreal Province, where the Mon tian s. s. is definitely the age equivalent of the Selandian s. s.with its upper exten sion above th e Selandian. Thi s superposition can be ob served especially inNorth Poland and to a lesser extend in Limburg Province, where the influence of the Paleocenetropical sea was decidedly younger than of boreal. A similar opinion of the Montian-Selandianrelation is held by NAKKADY (1957). Th e present investigations, showing the predominance ofSelandian microfauna below that of subtropical Montian microfauna, supports NAKKADY'S sup­positi on that "the Swedish-Dan ish Lower Paleocene is very similar to that of the type UpperDanian, would mean an earlier Terti ary transgression depositing the Scandinavian LowerPaleocene (Selandian), before the type Monti an , with a slight vertical extension to the base ofthe Thanetian" (1957, p. 433).

HOFKER (1956-1966) also is in agreement with the above view of the superpositionof the discussed stratigraphic un its of the lowermost Tertiary. According to the latter author,the Scandinavian Paleocene , as identified by BROTZEN (1948) in South Limburg, lies on theuppermost Maastrichtian (local horizon "Md") sediments, which are covered by "Tuffeaude Ciply" and "Calcaire de Mons", i. e. typical Montian layers, thus considering Selandians. s. as older than Montian s. s. On the other hand, an opposing view is held by ROSENKRANTZ(1963, MS) and BERGGREN (1962-1965).

The Paleocenc sedimenta tio n in the mainland of Europe, the geosynclinal region excluding,finished just above the uppermost Montian in several countries. Then fresh-water, lagoonarPaleocene (Thanetian) sediments appear. These are developed in France (ROUVILLOIS, 1960),Belgium and Holl and (HoFKER in 1966 described from there the so-called ,,s" Zone, referredby him to Thanetian). It is probable that the Kerteminde mar! of Denmark was also depositedPalaeoatologia Polon ica No . zo 3

34 KR YST YNA POZARYSKA & JANI NA SZCZEC HU RA

at that time . Thi s was suggested by BROTZEN (1959) who sta ted th at sediments, rich in Fora­minifera similar to that occurring in Middl e and U pper Paleocene sediments of England andWestern Europe , occur in the Selandian sediments of Denm ark.

The Tertiar y sedimentation in England began with the tr an sgression of the boreal Tha­netian sea. Its sedimenta tion, as well as the for aminiferal fauna it con tain s, can be comparedto that of Scandinavian Selandian . In the opinion of the present autho rs, the foraminiferalassemblage characteristic of Engli sh Thanetian, and its equivalent on th e mainland of Europe,are a continuation of the Scandinavian Selandian foraminiferal microfauna. HOFKER (1966,p. 327) considered that the "resembla nce between Th ane tian and Selandian is very meagre".HAYNES (1958), B ERGGREN (1964) and ROSENKR ANT Z (1924), however, suggested that Thanetiancorresponds in age to Selandian.

Due to the absence of Paleocene layer s younger than Montian in the Poli sh Lowlands,the present authors were unable to consider th e upper boundary of Paleocene. Thi s is also whythe present authors were not concerned with the Upper Paleocene and did not analyse substagesyounger than Thanetian. The Thanetian, although unknown in Poland , is discussed in the presentpaper on account of its similarity to Selandian.

The present authors ass igned the Danian and Montian to Paleocene, con sidering theMontian s. I. as at least partly an equivalent of Montian s. s. and Selandian s.s., this former ,however, with its upper extension above the Selandian. As alread y menti oned, HOFKER (1966)as well as other authors, e. g. B ROTZEN (1959), V . P. V ASILENKO (1961), JA NSIN (1960) and othersconsider the Paleocene as a syno nym from which the Tertiar y is dated, while ass igning Da­nian to the Cretaceous.

SY STEJYIA'TIC PAR_T

Family TEXTULARIIDAE EHRENBERG, 1838

Genu s SPIROPLECTAMMINA CUSHMAN, 1927

Spiroplectammina wilcoxensis CUSHMAN & PONTON, 1932

1932. Spiroplectammina wilcoxensis CUSHMAN & PONTON; J.A. CUSHMAN & G. M . PONTON, An Eocene foram iniferalfaun a.. . , p. 51, PI. 7, fig. 1.

1965. Spiroplectammina wilcoxensis CUSHMAN & PONTON; K. POZARYSKA, Foraminifera and biostratigraphy. . ., p. 50,PI. 3, fig. I (here additional synonymy included) .

Material. - About 20 specimens, most of them damaged on the apertural end .Dimension s, description and remarks are given in POZARYSKA'S paper (1965, p. 50).

Occurrence. - Not common in the Paleocene (Montian only) of Poli sh Lowlands and thePamietowo boring. Present in the Paleocene (Selandian) of Sweden , Denmark and in theEocene of America (D.S.A.).

Genus TEXTULARIA DEFRANCE, in DE BLAINVILLE, 1824

Textularia plummerae arkansasana CUSHMAN, 1951

1951. Textularia plummerae LALlCKER var. arkansasana CUSHMAN; J. A. CUSHMAN, Paleocene Fo raminifera. .. , p. 7,PI. 2, figs. 4-5.

1965. Textularia plummerae (LALlCKER); J. INDANS, Mikrofaunistische No rmalprofil.. ., PI. 1, fig. 4.1965. Textularia plummerae arkansasana CUSHMAN; K. POZARYSKA, Foraminifera and biostrat igraph y. .. , p. 51, PI.l ,

fig. 6a-b (here additional synonymy included).

Material. - Few specimens, damaged on both ends.Dimensions, description and remarks are given in POZARYSKA'S paper (1965).

Occurrence. - In Poland rare in the Paleocene, i.e. Danian and Montian (Selandianonly) of Poli sh Lowlands and the Pamietowo boring. Present in the lowermost Tertiary ofDen­mark, HoIIand, Belgium , Germany (Krefeld region) and in America (D. S. A.) , where it occursin the Paleocene (Midway), as weII as in somewhat younger stra ta.

36 KR YSTYNA POZAR YSKA & JANI NA SZCZECHURA

Family ATAXOPHRAGMlIDAE SCHWAGER, 1877

G enus ARENOBULIMINA CUSHMAN, 1927

Arenobulimina cuskleyae JENNI NGS, 1936

(Text -fig. 5)

1936. Arenobulimina cusk leyae JENNINGS; P. H. JENNINGS. A microfauna . . ., p, 13, PI. 1, fig. 8.1965. Arenobulimina cuskleyae JENNI NGS; K. POZARYSKA, Foraminifera and biostra tigraphy. . .. p. 56, PI. 4. fig. 8 (here

additional synonymy included) ,

Material. - About 30 specime ns, usually well pr eserved.Dimensions, description and remarks are given in POZARYSKA 'S paper (1965).

0.1mm..----;

Fig. 5Individual variation of Arenobulimina cusk leyae JENNINGS; Pamietowo boring, 243 m, Montian .

Variation concerns mainly the size and sha pe of tests. As is sho wn above (Text-fig. 5),there are probably meg alo spher ic and microspheric forms.

Occurrence. - In Poland not very co mmon in the Pale ocene, i. e. Dani an and M ontianin the Pamietowo, Sochaczew and Boryszew borings. Common in th e Swedi sh and DanishPaleocene (Seland ian) and in the Paleocene (M idway) of Am erica (D. S. A.).

G enu s VACUOVALVULINA HOFKER, 1966

Vacuovalvulina keijzeri (VAN BELLEN, 1946)

(PI. Ill , figs. 9-12)

1946. Marssonella k eij zeri VAN BELLEN ; R. c. VAN BELLEN, Foraminifera from the Middle Eoeene. . ., p, 30, PI. 2, figs. 3-5.1964. Conicovalvulina k eij zeri (VAN BELLEN); P. MARIE, Les facies du Mont ien .. ., p. I , fig. 3a-b.1966. Vacuovalvulina keyzeri (VAN BELLEN) ; J. HOFKER, Maestrieht ian , Danian and Paleocene. . ., p. 250, PI. 49, figs. 1-7.

Material. - About 20 specimens, ra ther well preser ved .Dimensions of two specimens (in mm) :

DiameterHeight

F. IX/150'320·18

F. IX/120·280'18

FORAMINIFERA FROM THE PALEOCENE OF POLAND 37

Description. - Test conical, slightly concave on ventral side, more elevated or less ondorsal side, rounded in outline. Chambers about 3 in number in the only visible whorl on theventral side, rapidly increasing in size as added, low and crescent on dorsal side. A distinctumbilical hollow on the ventral side, covered partially by a large valvular, slightly serrate tooth.Sutures flush with surface or slightly depressed, nearly invisible on ventral side, dorsally raised ,thickened, strongly curved. Surface except the ventral side fairly porous, somewhat rough.Aperture large, irregular, under the valvular tooth-lip-like, at the base of the last formedchamber.

Variation insignificant; it concerns mainly the size and shape of test. Specimens are ele­vated dorsally in different degree, with more developed or less developed valvular tooth (seePi. Ill, figs. 9-12).

Remarks. - Specimen described from Poland as Vacuovalvulina keijzeri represent un­doubtedly the species described by VAN BELLEN (1946) as Marssonella keijzeri from the Eocene(recte Montian) of Holland and cited and illustrated later by MARIE (1964) as Conicovalvulinakeijzeri from Ca1caire Pisolithique (Montian) of France, and by HOFKER (1966) as Vacuoval­vulina keijzeri from the Paleocene of Holland and Belgium. Our specimens, however, are generallyless conical dorsally and definitely sma ller.

Occurrence. - In Poland, not common in the Paleocene (Montian; in true Montianit is most common) in Pamietowo boring only. Known to be common in the Paleocene(Montian) of France (Ca1caire Pisolithique of Paris Basin), Belgium (Tuffeau de Ciply andCalcaire de Mon s, .Puits Artesien" in Mons) and Holland (Bunde).

Family LAGENIDAE CARPENTER, 1862

Genus ASTACOLUS DE MONTFORT, 1808

Astacolus arcuatus (PHlLlPPI, 1843)

1843. Marginulina arcuata PHILlPPI; R. A. PHlLlPPl, Beitrage zur Kcnntnis.. ., p. 5, PI. I, fig. 28.1965. Astacolus arcuatus (PHILlPPI) ; K. POZARYSKA, Foraminifera and biostratigraphy. 00 ' p. 77, PI. 10, fig. 6 a-b

(here additional synonymy included).

Material. - Some dozen specimens, in most cases well preserved.Dimension s, description and remarks are given in POZARYSKA'S paper (1965).

Variation insignificant; it applies mainly to the size and shape of test; specimens almostround or elongated in the outline occur. They have moreover sutural ornamentation, sometimesmore distinctly developed, sometimes less.

Occurrence. - In Poland, common in the Paleocene, i. e. Danian and Montian of PolishLowlands and Parnietowo boring. Known to be common in the Paleocene of Denmark (Se­landian) .

Astacolus gryi BRoTzEN, 1948

1948. Astaeolus gryi BROTZEN; F. BROTZEN, The Swedish Pal coc enc.. . , p, 44, PI. 7, fig. 6 a-c.1964. Astacolus gryi BROTZEN; K. POZARYSKA, On some Foraminifcra.v., p. 542, PI. 1, figs. 20, 21 ; PI. 3, figs. 1-25.1965. Astacolus gryi BROTZEN ; K. POZARYSKA, Foraminifera and bio stratigraphy... , p. 77, PI. 10, fig . I a-b.

38 KR YSTYNA POZARYSKA & JANINA SZCZECHURA

Material. - More than 30 specimens well preserved .Dimensions, description, a detailed illu strated variat ion and remarks are given III Po­

ZARYSKA'S papers (1964 , 1965).

Occurrence. - In Poland not very common in th e Paleoccne, i.e. Dani an a nd Montianof Polish Lowlands a nd th e Pamietowo boring. Present in the Paleocene (Selandian) of Denmark,Sweden and of U .S.S.R. (Russian Platform ).

Astacolus paleocenicus BROTZEN, 1948

(Tex t- fig. 6)

1948. Astacolus paleocenicus BROTZEN ; F. BROTZEN, Th e Swedi sh Paleocen e.. ., p. 43, PI. 7, fig. 7 a-b ; Text-fig. 8.1965. Astacolus paleocenicus BROTZEN ; K . POZARYSKA, Fo ra rninifera and biostra tigraphy... , p. 78, PI. 10, fig . 9.

Material. - A dozen or so specimens, sometimes damaged.Dimensions, description and remarks are given in POZARYSKA'S paper (1965).

1mm

2 4

Fig . 6Ont ogeny and individual var iat ion of Astacolus paleocenicus BROTZEN ; Pam ietowo boring, 207 m, Montian.

Variation rather significant. There are differences in size, shape and ornamentation oftests . The tests can be slim , either elongated or short and broad. The ornamentation can occurall over th e surface of the test , or can be find to its initial part, therefore the pattern differ s.Both the size and ornamentation are independent of the ontogenet ic development of test ,as is shown on the figure s above (see Text-fig. 6).

Occurrence. - In Pol and commo n in the Palcocene (Montian) of Polish Lowlands andPamietowo boring, where is less common. Known to be common in the Paleocene (Selandian)of Sweden and D enmark. It appears as rare in th e uppermost Danian.

FORAMINIFERA FROM THE PALEOCENE OF POLAND

Genus FRONDICULARIA DEFRANCE, ill D'ORBlGNY, 1826

Frondicularia biformis MARSSON, 1878

39

1878. Frondicuiaria biformis MARSSON; T. M ARSSON, Foramin ifera der weissen Sch reibkreide . . ., p . 137, PI. 2, fig. 17 c-d .1965. Frondicularia bifo rmis M ARSSON; K. POZARYSKA, Foraminifera a nd biost rat igraphy.. ., p . 82, PI. 11, fig. 1 (here

additional synonymy included) .

Material. - Some dozen specimens well preserved.Dimensions, description and variation are given in POZARYSKA'S papers (1957, 1965).

Remarks. - Within specimens from the Polish Montian there occur forms havinga different stage of development as shown on illustrations in POZARYSKA'S paper (1957, p. 139,fig. 33a-g).

Occurrence. - In Poland common in the uppermost Cretaceous, rarely met in the Pa­leocene, i.e . Danian and Montian . Kn own as common from the Upper Cretaceous of Europeand America.

Genu s MARGINULINA D'ORBIGNY, 1826

Marginulina costulata HOFKER, 1958

1951. Ma rginulina sp. B ; J. A. CUSHMAN, Paleocene Foramin ifera . .. , p . 18, PI. 5, figs. 32-33.

1957. Ma rginulina cf. armat a R EUSS; K . POZARYSKA, Lagenidae d u Cretace... , p, 105, PI. 11, fig. 13.1958. Marg inulina costulata HOFKER; J. HOFK ER, Forami ni fera from the Cretaceo us .. ., XXXIV, p. 66, Text-fig. lOa-b.1965. Marginulina costulata HOFKER; K. POZARYSKA, Foramini fera a nd biostratigraphy.. ., p . 68, PI. 9, figs. 1, 2.

Material. - Some dozen specimens well preserved.Dimensions, description, variation and remark s are given in POZARYSKA'S paper (1965).

Occurrence. - In Poland, common in the Paleocene (Montian) of Polish Lowlandsand Pamietowo borin g. Present in the Paleocene (Selandian) of Sweden and Denmark.

Genus ROBULUS DE MONTFORT, 1808

Robulus degolyeri (PLUMMER, 1926)

1926. Cristellaria degolyeri PLUMM ER ; H. J. PLUMMER, Fo ramini fera o f the Midway ... , p. 97, PI. 7, fig. 7.1964. Robulus degolyeri (PLUMMER); K. POZARYSKA, On so me Foraminifera .. ., p . 540, PI. 1, fig. 22.

1965. Robulus degolyeri (PLUMMER); K. POZARYSKA, Foraminifera and biostratigraphy. .. , p. 62, PI. 7, fig. 4 (here additionalsynonymy included).

Material. - About 30 specimens rather well preserved.Dimensions, description, variation and remark s are given 111 POZARYSKA'S paper (965).

Occurrence. - Jn Poland not common in the Paleocene (Montian only) ofPolish Lowlandsand the Parnietowo boring. Known to be common in the Paleocene (Midway) of America(U.S.A.).

40 KRYSTY NA POZARYSKA & JANINA SZCZECHURA

Robulus rancocasensis OLSSON, 1960

(Text-fig. 7)

1960 . Robulus rancocasensis O LSSON; K. R. OLSSOl', Fo raminifcra of the latest.. ., p. 10, PI. 2, figs. 5. 6.1965. Robulus rancocasensis OLSSON; K. POZARYSKA, F oram inifera a nd biostratigraphy . . ., p, 64, PI. 7, figs. 3a-b, Sa -c.

Material. - Several hund red specime ns well preserved.D imension s, descript ion and remarks are given in POZARYSKA'S paper (1965) .

O:l mrll~

Fig. 7

Robulus rancocasensis O LSSON: 1,2 megalospheric forms , 3. 4 microspheric forms; Pa rnietowo boring, 228 m , Montia n.

Variation insignifica nt ; it co ncern s mainly the size and sha pe of the tests. Tes ts canbe regularl y rounded in out line o r elo nga ted a t the a per tura l end . Microspher ic a nd mega­lospheric forms occur (see Text-fig. 7).

Occurrence. - In Pola nd , very common in the Pa leocene, i. e. in Dan ian and M ontian .Co mmon in the Pa leocene, i. e. Dan ian and Seland ian of Den mark and Sweden , and Paleocene(Midway) of Am erica (U.S.A.) .

Robulus turbinatus (PWMMER, 1926)

1926. Cristellaria turbinata PLUMMER; H. J. I' Lu MMER, For a minifcra of the Midway . .. , p. 93, PI. 7, fig. 4a-b.1965. Robulus turbinatus (PLUMMER); K . I'OZARYSKA. Forami nifera and biostr at igraphy . . ., p. 64, P I. 8, fig. 1a-b ( here

additional syno nymy included} ,

Material. - Abo ut 40 specime ns, in most ca ses well preserved .D imensions, descr iption and remark s are given in POZARYSKA'S paper (965).Va riation com pletely insignifica nt.

O ccurrence. - In Po land rat her common in the Paleocene , i. c. Da nia n and M ontia nof Poli sh Lo wland s a nd Pamietowo boring. K nown as common in Paleocenc, i.e. D ani anof Denmark an d M idway of America ( U.S .A .) .

Family POLYMORPH INIDAE D'ORBIGNY, 1839

Genu s GLO BULINA D'ORBIGNY, ill DE LA SAGRA. 1839

Globulina arenacea BROTZEN, 1948

1948. Globulina arenacea BROTZEN; F . BROTZEN, T he Swedish Pa leoccnc... , p . 48, Te xt- pl . 10, fig. 3a-d.1965. Globu/ina arenacea BROTZEN; K . POZARYSKA, Foraminifera and biostrat igraphy. .. , p. 85, PI. 22, fig. 5.

FORAMINIFERA FROM THE PALEOCENE OF POLAND 41

Material. - About 30 specimens well pre served.Dimensions and description are included in POZARYSKA'S paper ( 1965).

Variation is mainly observed in th e size a nd degree of tests ornamentation.

Occurrence. - In Poland , not common in Paleocene , i. e. Danian and Montian of PolishLowl ands and Parnietowo boring. Known as not common in Pale ocene (Selandian) of Sweden .

Globulina gibba gibba (D'ORBIGNY, 1826)

1826. Globulina gibba O'ORBlGNY; A. 0 'O RIlIGNY, Ann. Sci. Nat ., p. 266, modeles 63(f ide Cat. of Forarn., ELLIS & MESSINA).1960. Globulina gibba O'ORBlGNY; A. ROUVILLOIS, Le Thane ticn. .. , p. 59, PI. I, fig. 8.1965. Globulina gibba O'ORIlIGNY; K. POZARYSKA, Foramini fera and biost rati graphy... , p , 86, PI. 13, fig. 3 (here additional

synonymy included) .

Material. - Over one hundred specimens well pre served.Dimensions, description and remarks are given in POZARYSKA'S paper (1965) .

Variation conc erns the size and sha pe of th e test. Specimens can be more. ovat e or less,o r rounded in outl ine.

Occurrence. - In Poland, common in Paleocene, i. e. Danian and Montian in all stud iedlocalities. Kn own as co mmo n in the Lower Tertiary stra ta of Europe and America.

Globulina gibba multistriata n. subsp .

(PI. Ill , fig. 8)

unpu bl. Globulina gibba v. multistriata MARI E; P. MARIE, Foraminiferes du Calcai re Pisol ith iqu e. . ., (MS), PI. 11, fig. (i.

Ho lotypus: Specimen presented on PI. Ill , fig. 8 (F . IX /I ).Stratum typicum : Montian,Locus typicus: Pamie towo boring, depth 208·5 m (North Poland ).Derivatio nominis: Lat. multistriata = with many ribs, the name being suggested by P. MARIE (MS) .

Material. - Several specimens well preser ved .Dimen sions of an average specimen (in mm):

WidthHeight

F. IX/I0.330·37

Description. - Test almost sphaerica l, somewha t pointed on both ends, chambers few,3-4 in number, sutures flush with surface, quite invi sible , surface ornamented with numerousregular longitudinal rib s. Aperture terminal , radiate.

Variation indistinct, concerns mainly the sha pe of tests and number of surface rib s.

Remarks. - Our specimens in compari son with figured specimen of Globulina gibbainultistriata described by MARIE (MS) from the Montian of Belgium, are generally less compressedat both ends, having less sharp rib s. Because of the variability of our specimens it seems po ssiblethat there may be more th an one species .

Occurrence. - In Poland rare in Paleocene (Montian) of Pamietowo boring only. Presentin Montian (Calcaire de Mons) of Belgium, Holland and in France.

42 KRYSTYNA POZAR YSKA & .lANINA SZCZECH URA

Globulina cf. tu berculata O'ORBIGNY, 1846

(PI. III , fig. 6)

1846. Globulina tuberculata D'O RBIGNY; A. O'ORBlGNY, Foram inife rcs fossilc s.. ., p. 230 , PI. 13, figs. 21-22.1964. Globulina tuberculate D'O RBlGNY; J . HOFK ER, Foraminifcra from the Cretaceous. . ., LXXX, p. 117, fig. 13.

Material. - Several spec imens well preserved.D imensions of an average specimen (in mm) :

W idthHeight

F. IX!30-400·43

Description. - Te st somewha t ovat e in outlin e, mo re poi nted at the apertur a l end,ro und at th e ba se, chambers few, 3--4 in number, sutures and aperture not visib le, surfacecovered by numerous, irregu larly sca ttered, blunt a nd short knobs.

Variation not known beca use of th e sca rcity of material.

Remarks. -'- Our specime ns seem to be very simi lar to those described by O'ORBIGNYfrom the Miocene of Vienna Basin , but O'ORBlGNY'S figures are inadequate and too schema ticto be taken int o considerati on. Specimens ass igned to thi s species, described by HOFKER (1964)from the Paleocene of Holland (Curfs quar ry) , di ffer somewhat in compari son with those fromPoland in ornamentati on , being sca rcely a nd more regularl y tuberculated.

Occurrence. - In Poland ra re in the Paleocene (Montian ) in th e P arnietowo boringonly. Present in the Miocene of Vienna Basin.

Globulina sp.

(PI. Ill , fig. 7)

Material. - A few speci me ns well preserved.Dimensions of a n average speci me n (in mm):

F. IXj2Width 0· 33Height 0 ·33

Description. - Test about spherical, compressed a t both ends, chambers few, 3--4 innumber, sutures nearly invi sible, flush with surface. Surface covered by well developed inte r­ru pted ribs, thick and blunt , which here and the re pass into single tubercles.

Vari ation not known because of the sca rcity of mat erial.

Remar ks. - Our specime ns a re slightly similar to Globulina gibba var. striata EGGER,1857, from the Miocene of German y, d ifferin g however in having thicker and more regula rlyd ispersed rib s on th e surface of test.

O ccurrence. - In Poland rare in th e Paleocene (M o nt ian ) in the Parni etowo boring on ly.

Genus G UTTULINA O'ORBlGNY, ill OE LA SAGRA, 1839

Guttulina communis 0 'ORBlGNY. 1826

(Tex t-fig. 8)

1826. Guttulina communis D'O RBIGNY ; A. D'ORBlGNY, An n. Sci . Nat., no. 15, p. 266, PI. 12, figs. 1-4 (f ide Cat. of Foram.,

Et.us & MESSINA) .

FORAMINIFERA FROM THE PALEOCEN E OF POLAND 43

1960. Guttulina communis O'ORBIGNY; A. ROUVILLOIS, Le Than etien... , p, 58, PI. 1, fig. Sa-b.1965. Cuttulina communis O'ORBlGNY; K . POZARYSKA, Foraminifera and biostratigraphy.. ., p . 83, PI. 12, fig. I a-b (here

additional synonymy included) .

Material. - Several hundred specimens well preserved .Dimensions, description and remarks are given in POZARYSKA'S paper (1965).

Variation is mainly restricted to the size and shape of tests. Some examples are grvenbelow (see Text-fig. 8).

Fig. 8Individual variation of Cuttulina communis O'ORBIGNY; Parnietowo boring, 241 m, Montian .

Occurrence. - In Pol and very common in the uppermost Ma astrichtian, and Paleocenc,i. e. Danian and Montian in all studied localities of Polish Lowlands and in the Pamietowoboring. Known as common in Lower Tertiary sediments in Europe.

Guttulina problema D'ORBlGNY, 1826

(Text-fig. 9)

1826. Guttulina problema O'ORBIGNY ; A . O'ORBIGNY, Ann. Sci. Nat ., no. 14, p. 26 (fide Cat. of Foram., ELLIS & M ESSINA).1960. Cuttulina problema O'ORBIGNY; A. ROUVILLOIS, Le Than et ien ... , p. 58, PI. I , fig. 6a-b.1965. Guttulina problema O'ORBIGNY; K . POZARYSKA, Foraminifera and biostratigraphy... , p. 84, PI. 12, fig. 2a-h (here

additional synonymy included) .

2 3

0.1mm............---

F ig. 9Individual va riatio n of Guttulina problema O'ORBIGNY ; Pamietowo bo ring, 231 m , Montian .

Material. - Over two hundred specimens well preserved.Dimension s, description and remarks are given in POZARYSKA'S paper (1965).

Variation significant ; it concerns the size and the shape of the test. Some specimensvary in the arrangement of chambers, which are inflated to different degrees (see Text-fig. 9).

44 KRYSTYNA POZARYS KA & JANI NA SZCZECHURA

Occurrence. - In Poland very commo n in the Paleocene, i. e. D anian a nd Montian inall stud ied sa mples from Polish Lowlands a nd the Pam ietowo bori ng. Known to be commonfrom the Tertiary of Europe and America (V .S.A.).

Genus PSEUDOPOLYMORPHINA CUSHMAN & OZAWA, 1928

Pseudopolymorphina frondea (CUSHMAN, 1922)

1922. Bolivina fr om/ea CUSHMAN; J. A. CUSHMAN, The Foraminifera of the Mint Spr ing.. ., p, 126, PI. 29, fig . 3.1951. Polymorphina frondea (CUSHMAN); J. A. CUSHMAN, Palcocene Forarninifera of the Gu lf. v., p. 35, PI. 10, fig. 13.1965. Pseudopolymorphina frondea (CUSHMAN); K. POZARYSKA. Foraminifera and biostra tigraphy.. ., p. 87. PI. 12,

figs. 5-6 ( here addit ional synonymy included) .

Material. - Over 30 specimens well pr eserved.Dim ension s and description are given in POZARYSKA'S pap er (l965).

Variation insig nifican t; thi s is a n extremely unifor rn'[species.

Occurrence. - In Poland , rare in the Paleocene, i. e. D ani an a nd Montian of Poli shLowlands and th e Parni etowo boring. Kn own in America (V .S. A.) from the Paleocen e up tothe Oli gocene.

Pseudopolymorphina geijeri angusta BROTZEN, 1948

1948. Pseudopolym orphina geijeri angusta BROTZEN; F. BROTZEN, The Swedish Paleocene. .. , p. 52, Text-p I. 10, fig. 15 ;Text-fig. 12.

1965. Pseudopolymorphina geijeri angust« BR(HZEN; K . POZARYSKA, Foraminifera and biostratigraphy.. ., p. 88, PI. 12,fig. 8a-b .

Material. - Abo ut 20 specimens well prese rved.Dim ension s, descr iptio n a nd rema rk s are given in POZARYSKA'S paper (l965).Variation not kno wn becau se of the scarcity of mater ial.

Occurrence. - In Pola nd , not very comm on in the Paleocene, i. e. D anian a nd Montianof Pol ish Lowlands and the Pamietowo boring. Known from the Paleocene, i. e. Selandianof Sweden a nd U. S. S. R. (Do netz Basin ) and Mon tian of H olland (Bunde).

Pseudopolymorphina paleocenica BROTZEN, 1948

I94li. l'seudopolymorphina paleocenica BROTZEN; F. BROTZEN, The Swedish Palcocenc. . ., p. 50, Text-fig. 10 (fig. 13) ;Text-fig. I 1 (figs. 1-22).

1958. Pseudopolym orphina paleocenica BROTZEN; J . HAYNES, Certain smaller. .. , p. 11, PI. 4, figs. 7. 7b .1965. Pseudopolymo rphina paleocenica BROTZEN; K. POZARYSKA, Forarn inifcra and biostrati graph y.. ., p, 88, PI. 13,

fig. 6 (here addit ional synonymy included) .

Material. - Over 50 spec ime ns well preserved.Dimension s, descr iption and remarks are given 111 POZARYSKA'S paper (1965).

Variation insignificant; th e in fraspecific varia t ion observed within this species in Polandco rre spo nds to the ran ge var iatio n stud ied in deta il and illustrated by BROTZEN (1948) .

Occurrence. - In Poland, not very common in the Paleocene, i. e. Danian a nd Montianof Poli sh Lowlands and Pamieto wo boring. Known to be common in the Paleocene (Selandian)of Sweden and Denmark and in th e Paleocene of Holland, England and U. S. S. R. (Crimea).

FORAMINIFERA FROM THE PALEOCEN E OF POLAND

Genus PYRULINA D'ORBIGNY in DE LA SAGRA, 1839

Pyrulina fusiformis (ROEMER, 1838)

45

1838. Polymorphlna fusiformis ROEMER; F. A . ROEMER, Die Cephalopoden des Nord-Deu tsehen. .. , p. 386, PI. 3, fig. 37a-b,1930. Pyrulina [us ijormis (ROEMER); .I. A. CuSHMAN & Y. OZAWA, Monograph of the foraminifera l fami ly. . ., p. 55.

PI. 13, figs. 3-8.1965. Pyrulina fusifo rmis (ROEMER) ; K . POZARYSKA, Foraminifera and biostrat igraph y... , p. 86, P1. 13, fig. 4a-b (here

addit ional synonymy included ).

Material. - A dozen or so specimens well preserved.Dimensions and description are given in POZARYSKA'S paper (1965).

Variation insignificant ; th is is a highly distinctive species.

Remarks. - The identification of our specimens is based on BROTZEN'S (1948) andHAYNES'S (1958) papers, because of the inaccuracy of ROEMER'S figures. The specimens describedthere and included into Pyrulina fusiformis do not differ from those from Poland referred tothat species, Recent species described by CUSHMAN and OZAWA as P. fusifo rm is do not seemto belong to this species judging from their sha pe.

Occurrence. - In Poland , rare in the Paleocene (Montian only) of Polish Lowland sand Pamietowo boring . Present in lowerrnost Tertiary sediments in Europe and America(U .S.A.).

Genus SIGMOMORPHINA CUSHMAN & OZAWA, 1928

Sigmomorphina brotzeni HOFKER, 1957

1957. Sigmo morphina brot zeni HOFKER ; J. HOFKER, Fora minifera fro m the Cretaceo us. . ., XX II I, p. 18, Text-figs. 18-25.1965. Sigm omorphina brotzeni HOFKER ; K . POZARYSKA, Forami nifera and biostrat igraphy. . ., p. 89, PI. 12, fig. 3a-b

(here additional synonymy included).

Material. - About 70 specimens, some damaged.Dimensions, description and remark s are given in POZARYSKA'S paper (1965).

Variation significant according to the size and general shape of test. Stud ied in detailby HOFKER (1957, p. 18, Text-figs. 18-25).

Occurrence. - In Poland, not common in the Paleocene, i. e. rar e in Danian , more commonin Montian of Polish Lowlands and Pamietowo bor ing. Known to be common in the Paleocene ,i.e. Selandian of Sweden and Denmark and Montian of Holland , where it starts from uppermostMaastrichtian.

Sigmomorphina pseudoregularis CUSHMAN & THOMAS, 1930

1930. Sigmomorphina pseudoregularis CUSHMAN & THOMAS; J . A . CUSHMAN& Y. OZAWA. A mo nogra ph. . .• p, 125. PI. 32,fig. 8.

1948. S igmomorphina pseudoregularis CUSHMAN & THOMAS; F. BROTZEN, T he Swed ish Paleocene. . ., p. 54, Text-pl , 10.fig. 17.

1965. Sigmomorphina pseudoregularis CUSHMAN & THOMAS; K . POZARYSKA, Foraminifera and biostratigr aph y... , p. 89.

PI. 12, fig. 9 (here additional synonymy included) .

46 KR YSTYNA POZAR YSKA & JAN INA SZCZECHURA

Material. - - A dozen or so specimens well preserved.Dimensions, description , variation and remarks are given in POZARYSKA'S paper (1965).

Occurrence. - In Pol and , rare in the Paleocene (Mo ntian only) of Polish Lowlands andPamietowo boring. Common in Paleocene (Selandian) of Sweden and Denmark and in theEocene of North America.

Family GLANDULINIDAE Rsuss , 1860

Genu s GLANDULINA D'ORBIGNY, in DE LA SAGRA, 1839

Glandulina laevigata D'ORBIGNY, 1826

1826. Nodosaria (Glandulina) laevigata O'ORBIGNY ; A . O'O RBIGNY, T abl eau methodique... , p. 252 , PI. 10, figs. 1-3(fide Cat. of Foram., ELLIS & M ESSINA).

1846. Glandulina laevigata O'O RBIGNY; A.O'ORBIGNY, Forarni nif'eres fossiles... , p. 29, PI. I, figs. 4-5 .1965. Glandulina laevigata O'ORBIGNY; K. POZARYSKA, F oraminifera and biostratigraphy. .. , p, 9 1, PI. 22, fig. 7 (here

additional synonymy included) .

Material. - A do zen or so specimens, oft en dam aged .Dimension s, description and remarks are given. in POZARYSKA 'S pap er (1965).

Variation extremely insigni fican t ; this is a remarka bly individual spec ies.

Occurrence. - In Poland , rare in the Paleocene, i. e. Dani an and Montian of PolishLowlands and Parnietowo boring. Known to be common in the Upper Tertiary sedimentsin Europe.

Family TURRILINIDAE CUSHMAN, 1927

Genus PYRAMIDINA BROTZEN, 1940, nom. nud. , emend. BROTZEN, 1948

Pyramidina crassa BROTZEN, 1948

1948. Pyramidina crassa BROTZEN; F. BROTZEN, T he Swedish Paleocene. . ., p. 63, PI. 6, fig. 8.1965. Pyramidina crassa BROTZEN; K. POZARYSKA, Foramin ifera and biostratigraphy... , p. 99, PI. 15, fig. 5a-c.

Material. - About 30 specimens well pre served.Dimensions, description and remarks are given in POZARYSKA'S paper (1965) .

Variation insignificant ; it is rather a distinctive species. Some spec imens are twisted indifferent degrees.

Occurrence. - In Poland, not common in the Paleocene, i. e. Danian and Montian ofPoli sh Lowlands and Pamietowo boring. Common in the Paleocene (Selandian) of Swedenand Denmark and in U. S.S. R. (Russian Platform).

Pyramidina cuneata (BROTZEN, 1948)

1948. Angulogerina cuneata BROTZEN; F. BROTZEN, The Swed ish Pale ocene..., p. 64, PI. 6, fig. 10.1965. Angulogerina cuneata BROTZEN; K . POZARYSKA, Fora min ifera and biostratigraphy. .. , p. 99, PI. 15, fig. 7a-c.

Material. - Over two hundred specimens well preserved.Dimensions and description are given in POZARYSKA'S paper (1965) .

FORAMINIFERA FROM THE PALEOCENE OF POLAND 47

Variation insignific ant; it applies generally to th e size and development of marginalkeels; th ey can be more distin ct or less. M oreover , pa rt of th e specimens is somewhat twi sted.

Remarks. - According to BROTZEN (1948), th e forms belonging to Angulogerina genusa lways have an aperture on the neck. Illustr ated holotype of Angulogerina cuneata, speciesdescribed by BROTZEN (1948) from th e Paleocene of Sweden, is without a neck as a ll specimensoccurring in th e Poli sh specimens of Montian age . For th at reaso n th e mentioned species isreferred here to Pyramidina genus.

Occurrence. - In P oland, very common in the Paleocene, i.e. Danian and Montianof Polish Lowlands and Parni etowo boring. Known to be common in th e Paleocene (Selandian)of Sweden and Denmark.

Family BOLIVINITIDAE CUSHMAN, 1927

Genus BOLIVINA O'ORBlGNY, 1839

Bolivina oedumi BROTZEN, 1948

1948. Bolivina iidumi BR OTZEN; F. B ROTZEN, The Swedish Paleocene..., p. 65, P I. 9, figs. 3-4.1965. Bolivinita oedumi (BROTZEN) ; K. POZARYSKA, Foraminifcra and biostrat igraph y... , p. 101, PI. 15, fig. 13.

1965. Bolivinotdes oedumi (BROTZEN); B. M CGo w RAN, Two Palcocene.. .. p. 41, PI. I , fig. 6.

Material. - Several specimens well preser ved .Dimensions and de scription are given in POZARYSKA'S paper (1965) .Variation not known due to the sca rcity of material.

Occurrence. - In Poland, extremely rare in the Paleocene , i. e. Danian and Montianof Polish Lowlands and Pamietowo boring. Known to be rare in the Maastrichtian of Germanyand Holland, in th e Paleocene of Sweden (Selandian), and in Paleocene of Australia (WesternVictoria). Probably present in the Paleocene, i.e. Thanetian of England.

Family BULIMINIDAE JONES, 1875

Genus BULIMINA O'ORBIGNY, 1846

Bulimina ovata D'ORBlGNY, 1846

(Text-fig. 10)

1846. Bulimina ovata O'ORBIGNY ; A. O'ORDlGNY, Fo raminiferes fossiles. .. , p. 185, PI. 11, figs. 13-14.1954. Praeglobulimina ovata (O' O RBIGNY) ; J. H AYNES, Taxonomic position ... , p, 190, Text-figs. 9-12, 17-19.1965. Bulimina ovata O'ORBIGNY; K. P OZARYSKA, Foraminifera and biostrat igraphy ... , p. 98, P I. 15, fig. 10 (here additional

synonymy included).

Material. - About two hundred specimens well preserv ed.Dimensions, de scription and remarks are given in POZARYSKA'S paper (1965).

Variation insignificant; it is mainly found in the size, width/length ratio of tests and ar-rangement of chambers. There are both microspheric and megalospheric form s (see Text-fig. 10).

4R KRYST YNA POZARYSKA & JANINA SZCZECHURA

Fig. 10Bulimina ovata D'ORBlGNY ; J, 2, 5 megalos pheric fo rms, 3,4 m icro spheric forms ; Pamietowo boring, 257 m, Montian .

Occurrence. - In Poland common in the Paleocene, i. e. Danian and Montian everywhere111 Poli sh Lowlands and the Pamietowo boring. Known from th e upperm ost Maastrichtianup to the Upper Eocene in Europe and America (V .S. A.).

Bulimina paleocenica BROTZEN, 1948

1948. Bu/imina (Reussella) paleocenica BROTZEN; F. BROTZEN, T he Swe dish Pal eoccne ... , p . 6 1, PI. 6, figs. 5-6.1958. Bulimina paleocenica BROTZEN; J. HAYNES, Certa in sma ller .. ., p. 74, PI. 16, fig. 17.1965. BuIimina paleocenica BROTZEN; K. POZARYSKA, Foramini fera and biostratigraphy.. ., p, 98, PI. 15, fig. 4 (here

additional synonymy included) .

Material. - About one hundred specimens well preserved.Dimension s, description and rem arks are given in POZARYSKA'S pap er (1965).

Variation rather insignificant ; it concerns mainly th e length/width ratio of test a nd tomargins rounded or angled . Some specimens are slightly tw isted. Studied in detail by BROTZEN(1948 , pp. 61-62).

Occurrence. - In Poland, common in th e Paleocene , i. e. Danian an d M ontian sediments.Present in the Paleocene, i. e. Danian a nd Selandian of Denmark, Sweden , Paleocene ofHolland , England (Thanetian) and sout h regions of U . S. S. R. (Russian Platform).

Family UVIGERINIDAE HAECKEL, 1894

Genus ANGULOGERINA CUSHMAN, 1927

Angulogerina europaea CUSHMAN & EDWARDS, 1937

(PI. I, figs . 1-7)

1937. Angulogerina europaea C USHMAN & EDWARDS; J. A. CUSHMAN & P. G. EOWAROS, N ote on the ea rly.. ., p, 61,

PI. 8, figs. 17a-b, 18.1948. Angulogerina europaea CUSHMAN & EOWAROS ; F. BROTZEN, Th e Swedish Paleocene.. ., p, 64, PI. 6, fig. 9.

FO RAMINIFER A PAL EOCENE F ROM T HE OF PO LA ND 49

1966. Reussella enropaea (CUSHMAN & EOWARDS) ; J . H OFKER. Maastrichtian, Dan ian and Paleocene . .. , p. 242. PI. 49,fig. 58.

1962. Angulogerina europaca C USHMAN & E OWA RDS; M . E . S CHMID . D ie For am iniferenfauna des Bruderndorfer .. ..p. 334. PI. 4. fig. 7.

Material. - A dozen or so specimen s. in most cases well pre ser ved.Dimension s of 3 specimens (in mm) :

F. IX/6Height of test O'36Maximu m width 0·16

F. IX j70·400·20

F. IX/90·300·15

Longest diame terShortest diameterHeight of test

Description. - Test triserial, tapering, almo st tri angular in a horizontal section. Chambersgrowing rapidly in size as added, inflat ed or flattened , overh an ging each other, projecting moreor less on the margins, with excavations developed to different degrees at their base, where theyare bordered sometimes by a slight ridge. Sutures depressed , curved or sickled, sometimesstrongly sigmoid . Surface generally rough , porous. Ap erture terminal , basal , loop-shaped.

Variation . significa nt; it concern s the size and shape of the tests, which are more twistedor less, having angles more rounded or less rounded. Moreover the chambers are overhangingeach ot her to different degrees, having excavations at their base differently developed. Theparticular chambers are not always bordered on their lower part by dist inct ridges.

Remarks. - Our specime ns are very similar to th e holotype of this species described byCUSHMAN and EDWARDS from the Montian of Pari s Basin. The Swedish specimens referredby BRoTzEN to th is species slight ly differ from the hol otype, as well as from Polish spec imens,in having well develop ed three longitudinal ridges along th e most external parts of the test.It is probably relat ed with Angulogerina muralis (TERQuEM, 1882), known from the Eoceneof Franco-Belgian Basin.

Occurrence. - In Poland, not common in th e Paleocene (Montian) in Pamietowo boringonly. Known to be very rare in Paleocene, i.e. Danian and Selandian of Sweden, Montianand Thanetian of Paris Basin, Montian of Holland (Bunde boring), Thanetian of England.

Family DISCORBIDAE EHRENBERG, 1838

Genus DISCORBIS LAMARCK, 1804

Discorbis quadrata (TERQUEM, 1882)

(PI. 11, fig. 3)

1882. Rosalina quadrata T ERQUEM; O. T ERQUEM, Les Foraminiferes de I'Eocene.. .. p. 98. PI. 10. fig. 12.1949. Discorbis quadrata (TERQUEM) ; Y. L E C ALVEZ. Revision. .. . p. 25, PI. 2. figs. 21-23.1961. Discorbis quadrata (TERQUEM); J . P. H. K AASSCHIETER, Fo rami nifera of the Eocene... , p. 208, PI. 10. fig. 17a-c.

Material. - A dozen or so spec imens well preserved.Dimensions of an average specimens (in mm) :

F.IX/II0·350·300'15

Pa lae o n to logia Po lonica N u. 20 4

50 KR YSTYNA POZARYSKA & JANINA SZCZECHURA

Description. - Test plane -co nvex, outline rounded or slightly lobulate, peripheral marginacute, bluntly keeled, ventral side slightly con cave, with a sma ll umbilical depression, spiralside con vex. Ch ambers flattened, low, 4-6 in number in the only visible last whorl on ventralside . Sutures sick led, tr an sparent, flush with sur face or sometimes a little raised on dorsal sidedepressed on ventral one. Surface smooth , indistinctly finely perforated on both sides. Ap ei ture,a low slit-like interio marginal arch or a large, regular sutural opening at the ba se of the lastchamber, near the umbilicu s.

Variation insignifican t, mainly concerning the genera l shape and cha racter of sutures.Spiral side can be more co nvex or less.

Remarks. - Our specimens are, on one hand, nearly the same as those described byLE CALVEZ (1949) who revised the TERQu EM's species from the Lutetian of Paris Basin and,on the other hand, as those described by KAASSCHlETER (1961) from the Eocene of Belgium.Specimen s from the Montian of Poland generally have more elongated chambers and moresickled sutures, mo reover they have a more finely po rous surface in comparison with specimensfrom Belgium. In the present aut ho rs' opinion this species is very clo se to Discorbis ubiquaLE CALVEZ (1949, p . 23, PI. 2, figs. 27-29), as was stated by KAASSCHlETER (1961 , p . 209).At first sight Discorbis quadrata (TERQUEM) is very similar to Trochammina sp . (HOFKER, 1962,p. 10), its walls, however , are are naceous, without pores.

Occurrence. - In Poland , not commo n in the Paleocene (M ontian) of the Pamietowoboring. Present in the Paleocene (M ontian) of Belgium ("Puits Artesien" in Mons). Known tobe very rare in the Eocene of France and Belgium .

Genus EPISTOMINELLA HUSEZIMA & MARUHASI, 1944

Epistominella cr. limburgensis (VISSER, 1951)

(Text-fig. 11)

1946. Eponides minimus CUSHMAN; R. C. VAN BELLEN, Fora minifera .. ., p. 58, PI. 7, figs. 10-12 (non Eponides minimaCUSHMAN 1933, in CCL 9, p. 17, PI. 2, fig. 8).

71951. Pseudoparrella limburgensis VISSER; A. M. VISSER, Monograph on the Foraminifera .. ., p. 278, PI. 7, fig. 10.1960. Eponides acutus ROUVILLOIS; A. ROUVILLOIS, Le Thanetien.. ., p. 68, PI. 2, fig. 33a-c.1961. Pseudoparrrella limburgensis VISSER ; J. HOFKER, Foraminifcra fro m the Cretaceous , LIIl , p. 67, figs. 9-10.1965. Pseudoparrella limburgensis VISSER; K . POZARYSKA, Foraminifera and biostratigraphy , p. 113, PI. 16, fig. 5a-c.

Material. - One hundred specimens well preserved.Dimensions and descript ion are given in POZARYSKA'S pap er (1965).

Variation insignificant ; it app lies mainly to the genera l shape of tests and to the positionof aperture, as is seen on figures below (Text-fig. 11).

Remarks. - Polish specimens are very similar to those distinguished by HOFKER (1961)as megalo spheric form s of Pseudoparrella limburgensis VISSER (1951) . In HOFKER'S opinion,P. limburgensis (recte Epistominella limburgensis) is represented by two forms, A and B, outof which only megalospheric for m probabl y occurs in the Montian of Poland. Both formsdiffer greatly and deviate, however, from the holotype of P. limburgensis illustrated by VISSER(holotype of the discussed species was designated by HOFKER in 1961 as a microspheric form) ;so it is qu ite possible that they represent a different species, not con specific with Pseudoparrellalimburgensis s.s. For that reason, the presence of representatives of Epistominella limburgensisin the Montian of Poland is confirmed only tentatively.

FORAMINIFERA FROM THE PALEOCENE OF POLAND 51

Poli sh specimens are the same as those descr ibed and illustrat ed by ROUVILLOIS (1960)as Eponides acutus, and by VAN BELLEN (1946) as E. minimus CUSHMAN, but d iffer considerablyfrom th ose described by CUSHMAN (1933) as E. minima . T ha t is why specimens de~cribed byVAN BELLEN and ROUVI LLOIS are put by the present writers int o the synonymy of Epistomi­nella cf. liinburgensis VISSER. The generic name of this species has been adapted after Treatiseof Invertebrat e Paleon tology (C , 2, 1964), where LOEBLlCH and TAPPAN revised generic nameof Pseudoparrella CUSHMAN & TEN DAM, 1948, changing it into Epistominella HUSEZlMA & MA­RUHASI, 1944.

0.1 mm.------<

2

4

Longest d iameterSho rtes t diameterHeight of test

Fig. I1Individua l variation of Epistominel la cf. limburgensis (VISSEI); Pam ietowo bo ring, 257 Ill, Montian.

Occurrence. - In Poland, not very common in th e Paleocene, i. e. Danian and Montianof Polish Lowlands and Pam ietowo bo ring. Known to be common in th e Paleocene, i. e. Danianand Selandian of Denmark, Mont ian of Belgium ("Puits Artesien" in Mons) and Holland(Geulhem, Bunde). Present in the Paleocene of U .S. S. R . (Russian Platform) and in the Pa­leocene (Thanetian) of France (Paris Basin) and Eng land.

Genus ROSALINA D'ORBIGNY, 1826

Rosalina brotzeni HOFKER, \96\

(PI. H, fig. 1)

1961. Rosalina brotzeni HOf KER ; J. HOFKER, Foraminifera from the Cretaceous... , LIB, p. 63, fig. 3a-c.1962. Rosalina brotzeni HOf KER ; J. HOFKER, Correlation of tuff cha lk , p, 1081, Text-fig. 25c.1965. Conorbina brotzeni (HOFKER) ; B. MCGOWRAN, Two Paleocene , p. 45, PI. 2, fig. 4.1966. Rosalina brotzeni HOFKER; J. HOFKER, Maestrichtian , Dania n a nd Paleocen e.. ., p. 225, PI. 46 , fig. 138.

Material. - A few specimens, rather well prese rved .Dimensions of an ave rage specimen (in mm):

F. IXjl 60·200·160· 11

Description. - Test small, tr ochospiral , more convex on dorsal side than on ventralon e, nearl y rounded , somewhat lobulate in outline, periphery slightly acute with roll-likethicken ings along the edge. Chambers crescent on dorsal side , inflat ed, indistinctly arrangedin 2-3 who rls, the last containing abo ut 5 chambers visible on the ventral side. Sutures radial,flush with sur face and curved on the dorsal side, somewhat depressed , enlarging at the open,

4·

52 KRYSTYNA POZARYSKA & JANI NA SZCZECHURA

Longest diameterShortest diameterHeight of test

cross-like umbilicus on ventral side. Surface rather smooth, glassy, finely porous. Apertureslit-like, obscured at the base of the last chamber, between the periphery and the umbilicus.

Variation insignificant, concerning mainly the general appearance of tests.

Remarks. - Specimens referred to Rosalina brotzeni HOFKER from Poland are verysimilar to those described under the same specific name by HOFKER (1961) from the holes inthe hard ground of Curfs quarry (Holland). Specimens from Poland are smaller, somewhatmore convex on the ventral side and less rounded, with a roll-like keel on their periphery.

Occurrence. - In Poland, rare in the Paleocene (Montian) in Pamietowo boring only.Known from Paleocene of Holland (Montian) and of Australia.

Rosalina koeneni BROTZEN, 1948

(PI. 11, fig. 2)

1948. Rosalina koeneni BROTZEN; F . BROTZEN, The Swedish Paleocene , p. 73, PI. 9, fig. 11.1956. Rosalina koeneni BROTZEN; J. H AYNES, Certain smaller British , p. 91, PI. 16, fig. 12a-d.1961. Rosalina koeneni BROTZEN; J. HOFKER, F or aminifera from the Cretaceous... , UII, p. 63, Fig. 2.1962. Rosalina koeneni BROTZEN; J. HOFKER, Correlation..., Text-fig. 25B.1966. Rosalina koeneni BROTZEN; J. HOFK ER, Maestrichtian, Danian and Paleocene Foraminifera.. ., p . 225, PI. 54,

fig. 82a-c.

Material. - Several specimens well preserved.Dimensions of an average specimen (in mm):

F. IX/170-460·400·15

Description. - Test small, trochospiral, elevated on dorsal side, concave on ventralone, rounded or somewhat ovate, not lobulate in outline, periphery sharply acute with welldeveloped sharp keel. Chambers slightly inflated, arranged into three whorls; the last, con­sisting of 6-7 chambers, is seen on the ventral side. Sutures strongly curved and flush withsurface on dorsal side , slightly depressed and nearly radial on ventral side, pa ssing into a star­shaped large umbilicus; an umbilical hollow filled up by a small but distinct, elevated plug.Surface smooth, finely porous. Aperture slit-like near the umbilicus, under a lip at the baseof the last formed chamber.

Variation not known, due to the scarcity of material.

Remarks. - Our specimens do not differ either from those described from Sweden byBROTZEN (1948) or from those described by HOFKER (1961, 1962, 1966) from Holland.

Occurrence. -In Poland rare, in the Paleocene (Montian) of Parnietowo boring only.Knownto be scarce in the Paleocene, i. e. Selandian of Sweden and Denmark, Montian ofHolland, Thanetian of England.

Rosalina selandiana n. sp.

(PI. 11, fig. 4; Text-fig. 12)

1966. Neoconorbina (Rosalina ) parisiensis (D'ORBIRNY); J. HOFKER, Maestrichtian, Danian and Paleocene. .. , p. 226,PI. 42, fig. 67) (non Rosalina parisiensis D'ORBIGNY, 1826, p. 105, No. 5 - fide Cat. of Foram., ELLIS & MESSINA).

Longest diameterSho rtest diam eterHeight of test

FORAMINIFERA FROM TH E PALEOCE NE OF POLAND

Holotypus: Specimen presented on PI. IT, fig. 4 (F. IX/18).Stratum typicum : Montian.Locus typicus: Nasil6w outcro p, 3 m above the Upper Maastrichti an hard gro und.Derivatio nominis: Lat. selandiana - fro m Selandian , Scandinavian Lower Paleocene.

Material. - A do zen or so specimen well pre served .Dimensions of an average specimen (holotype) in mm :

F. IX/I t'0·230·200·10

53

Description. - Test small, trochospiral, con vex on dorsal side, flat or even slightly con­cave on ventral side, outline rounded and insignificantly lobulate, periphery acute with a weaklydeveloped keel. Chambers crescent on the dorsal side, coiled into over two whorls indistinctlyseen on the dorsal side, 5-6 in number in the last whorl ; sutures stro ngly cur ved, limbate,flush with surface on dor sal side, not visible on ventral side. Large umbilical dep ression coveredby irregular ripple s radially scattered . Surface smooth, finely pitted, ap erture obscured cannotbe observed.

Variation insignificant, concern ing mainly the development of ripp les in the middle ofventral side, as is shown below (see Text-fig. 12).

Remarks. - Our specimens of Rosalina selandiana n. sp. differ from specimens of so fardescribed species of Rosalina mainly in their orna mentation. Our specimens do not seemto differ from specimens described by HOFKER (1966) as Neoconorbina (Rosalina) parisiensis(D'ORBIGNY) from Holland. Those last ones, howe ver, differ con siderably from the figuredspecimen of Rosalina parisiensis D'ORB. (1826) describ ed from Paris Basin, so in the presentpaper they are included into the here erected new species R. selandiana.

I a 2a 2b

Fig . 12Indiv idu al variation of Rosalina selandiana n. sp. ; Pam ieto wo boring, 210·7 m, Montian.

Occurrence. - In Poland, not common in the Paleocene (Montian) of Polish Lowlandsand Pamietowo boring, also in Paleocene of Holland. Probably present in the Paleocene,i. e. ,Thanetian of France (Paris Basin) and of England.

Rosalina ystadiensis BROTZEN, 1948

(PI. XVI, fig. 7)

1948. Rosalina ystadiensis BROTZEN; F . BROTZEN, Th e Swedi sh Paleocene , p. 72, PI. 9, fig. 9.1956. Rosalina cf. ys tadiensis BROTZEN; J. HAYNES, Certain sma ller British , p. 92, PI. 16, fig. 17.1961. Rosa/ina ys tadiensis BROTZEN ; J. HOFKER, Foraminifera from the Cretace ous... , LIH, p. 63, Text-fig. la-c.

54 KRYSTYNA PO ZAR YSKA & JA N1NA SZC ZEC H U RA

1962. Rosalina ystadiensis BROTZEN; J. HOFKER; Co rrela tion of the tuft' chalk.. ., Text-fig. 25A .1966. Rosalina ys tadiensis BROTZEN; J. HOFKER. Maestrichtian, Danian and Pa1eocene.. ., p, 225. PI. 45, fig. 137.

Material. - A single specimen well preserved.Dimension s of one specimen (in mm) :

Longest diam eterShortest diamete rHeight of test

F. IXjl 90·250·200·20

Longest diameterShorte st diameterHeight of test

Description. - Test very small, somewhat coni cal, spiral side elevated, umbilical sideconcave, involute. Outline rounded, ind istinctly lobulate, peripheral margin abruptly keeledand transparent, with radi al marks. Chambers crescent on do rsa l side, arranged in at least3 whorl s, the outer one contain ing 5 chambers; on the ventral side chambers of last whorlvisible, being sepa ra ted by radial, depressed sutures. On the dor sal side sutures transparen tflush with sur face. Umbilical depression sha llow but distinct, connected with slit-like aperturebelow the last chamber and penetrating into sutures within the umbilicus. Wall surface slightlyperforated, more on ventral side.

Variation not known due to the scarcity of materia l.

Remarks. - Our specimen differs from the illustrat ed holotype of Rosalina ys tadiensisBROTZEN, 1948 from Sweden in not having any tubercles inside th e umbi lical depression .

Occurrence. - In Poland, a single specimen in the Pa leocene (Mo ntian) of Pamietowoboring. Very rarely fou nd in the PaIcocene of Sweden (Selandian) and Holland (Montian).Probably also present in Englan d (Th anet ian).

Genus ROTORBINELLA B AND Y, 1944

Rotorbinella mari ci (VAN B ELLEN , 1946) 2

(PI. IV. figs. 1-3 ; PI. XVII. figs. 1·4)

1946. Parrella mariei VAN BELLEN; R. C. VAN BELLEN, Forarninifcra From the Middle Eocene.. ., p. 70. PI. 10, figs. 1-6,?1957. Rotorbinella mariei (VAN BELLEN); J. HOFKER. Fora rninifc ra From the Cretaceo us... , XXX, p. 123, fig. 2, non

fig. 1.?1959. Rotorbinella mariei (VAN BELLE1-I) ; J. HOFKER, Les Fo ra rn iniferes des craies tuffoldes, p . 293, figs. 110-112.

1964. Poi aryskaia mariei (VAN BELLEN) ; P. MARIE, Les Iacies du Mont ien.. .. p. 1098, PI. 2, fig. 5a-c.1966. Rotorbinella mar iei (VAN BELLE1-I) ; J. HorKER, Macstrich t ian, Danian and Paleocene.. .. p. 257, PI. 54, figs. 91-93.

?1966 . Rotorbinella bundensis (VA1-I BELLEN) ; J. HOFKER, Ibid.• p. 258, PI. 54, figs. 88-90 (non Rota/ia bundensisVAN BELLEN; R . C. VAN BELLEN. 1946, Foraminifera ..., p , 61, PI. 8, figs . 1-4),

Material. - Two dozen specimens, in most cases well pre served.Dimension s of an average specimen (in mm):

F. IXj470·17

0·150·09

• The genus Pozarysk aia instead of Rotorbinella was proposed by MARIE (1964) as nom ina nuda. Th at is why weuse here the earlier generic name For the discussed species.

FORAMINIFERA FROM THE PALEOCENE OF POLAND 55

Longest diameterShortest diameter

Height of te st

Description. - Test pIano-convex, dorsal side convex, ventral side flat or even slightlyconcave, outline rounded, fairly lobulate, periphery broadly rounded, generally with a keel-likelist formed by a marginal prolongation of sutural thickening on the dorsal side. Chambersslightly inflated on both sides, arranged into at least two poorly visible whorls, about sevenin number in the last whorl, very slowly increasing in size as added. Sutures indi stinct, gentlydepressed on ventral side in contrast with rai sed, limbate, rad ial sutures on dorsal side. Um­bilicus a deep, rather large hollow, filled up with numerous, regular, sma ll bosses. Surface roughto spiny on dorsal side, smooth only at the periphery of ventral side. Ap erture a large shortopening at the base of the last chamber, between the umbilicus and periphery on the ventralside.

Variation significant; it applies mainly to the ornamentation of tests on the dorsal side,where spines are not always well developed and sutures not raised , in which case the peripheralkeel is also lacking. Small bosses on the ventral side vary in number and arrangement.

Remarks. - Our specimens differ only in size from the illu strated specimens describedby VAN BELLEN (1946) as Parrella mariei and from MARIE'S (1964) one s assigned by him toPozaryskaia mariei, both from the Montian of Holland. It seems that specimens, describedby HOFKER (1957, 1959 not 1966) from the Maastrichtian ("Md" horizon) of Limburg, assignedto Rotorbinella mariei, belong to another species, on the ba se to their general habitus andornamentation. According to the pre sent authors' opinion , Rotorbinella mariei (VAN BELLEN)is probably related to Discorbis bundensis (recte Rotorbin ella bundensisi (VAN BELLEN, 1946,p. 47, PI. 4, figs. 19-24) (see a lso Rotorbinella bundensis [VAN BELLEN] HOFKER, 1966), fromwhich it differs in being coarsely pitted on the dorsal side between sutures ; sometimes evenit has weakly developed protuberances, asvis seen in ' comparative material from Holland(Bunde) (see PI. XVII, figs. 1-4).

Occurrence. - In Pol and not common in the Paleocene (Montian) of Parnietowo boringonly. Common in the Paleocene (Montian) of Holland (Bunde), Belgium (.,Puits Artesien" inMons) and France.

Rotorbinella montiana n. sp .

(PI. VI, figs. 5-6)

1946. Discorbis corrugata (C USHMAN & BERMUDEZ); R . c. VAN BELLEN, Fora minifera from the Middle Eocene.. ., p. 49,PI. 5, figs. 7-13 (non Anomalina corrugate CUSHMAN & BERMUDEZ; J. A. CUSHMAN & P. .I. BERMUDEZ (1937), p. 27,PI. 2, figs. 57-59).

1965. Discorbis corrugata (CUSHMAN & BERMUDEZ); J. INDANS, Mikrofaunistisches Normalprofil. .. , p. 13.1966. Rotorbinella corrugate (CUSHMAN & BERMUDEZ); .I. HOFKER, Maestrichtian, Danian and Paleocen e. 00 ' p. 258,

PI. 54, figs. 85-87.

Holotypus: Specimen presentes o n PI. VI , fig. 5 (F. IX/50 ).Paratypu s: Speci men presented on PI. VI , fig. 6 (F. IX/51) .Stratum typicum : Montian.Locus typicus: Pam ietowo boring, depth 210·7 m (North Poland).Derivatio nominis: Lat. montiana - from Mon tian .

Material. - About 30 specimens damaged.Dimensions of an average specimen (holotype) 111 mm:

F. IX/500·25

0'230·12

56 KR YSTYNA POZAR YSK A & JANINA SZCZECHURA

Description. - Test piano-con vex, dorsal side slightly convex, ventra l side flat o r evengently concave. Outline nearl y round, not lobulat e except between th e last chambers, peripherybluntly acute, roll -like. Ch ambers globularly inflated on dorsal side, rapidly increasing in sizeas added, arran ged in two whorl s, the last consist ing of about 7 cha mbers visible. Chambersfused on the vent ra l side . Sutures ind istinct, rather de pressed on dorsal side, flush with surface,obscured on ventra l side . Umbilicus large, with severa l kno bs, rather sha llow, irregularly de­veloped, filled with one or severa l plugs, which a re pr on ounced sometimes more, sometimes less.Numerous protuberances and severa l pits are develop ed aro und th e umbilicus. A smooth,rather wide border occurs alon g th e periphery on the ventra l side . Aperture a slit-like betweenthe periphery and the umbilicus.

Variation rather significant ; it applies mainly to the ornamenta tion of ventral side. Tu ­bercles are not always numerous and are developed to different degrees.

Remarks. - Our specimens in compa rison wit h illustrat ed French specimens (in MARIE'Smanuscript as Rotorbinoides montis n. sp.) seem to represent the same species, althought th ePolish specimens have less who rls an d are not ornamented at the periphery on the vent ra l side;moreover their peripheral bo rder is much wider. Our specimens are of the same appearanceas those described by VAN BELLEN (1946) as Discorbis corrugata, found by the present authorsin samples from the Montian of Holland (Bunde), where they have, however , a greater variation .Poli sh specimens are much smaller in compari son with Dutch on es. It is fairly sure that spe­cimens described by VAN BELLEN as D. corrugata do not differ from th ose of Rotorbinellamontiana n. sp . Specimens described by CUSHMAN and BERMUDEZ (1937) as Anomalina corrugatafrom the Eocene of Cuba are of quite different shape and ornamentat ion .

Occurrence. - In Pol and not commo n in the Paleocene (Montian) of Parni eto wo boringonly. Common in Paleocene (Montian) of Holland (Bunde and Beatrix bo rings), Belgium("Puits Arte sien" in M ons), F rance (Calcaire Pisolithique of Paris Basin) and Germany (Krefeldregion). Rotorbinella montiana n. sp. seems to be a goo d index fossil for true Montian.

Rotorbinella papillata n. s p .

(PI. VI, fig. 11 )

Holotypus: Specimen presented on PI. VI. fig. 11 (F . IX/52).St ratum typicum: Mon tian.Locus typicus: Pamietowo boring, dept h 210·7 m (North Poland).Derivatio nominis : Lat. papillat a - wit h a ro ugh, coarsely pitted surface.

Material. - Three speci mens well preser ved .D imensions of one spec imen, holotype (in mm) :

Longest diameterShortest diameterHeight of test

F. IX/520·130·\00·05

Description. - Te st sma ll, planispiral, ventra l side nearly flat o r even slightly co ncave,dorsal side somewha t convex, round in outline, lobulat e, periphery truncat ed , forming a roll-likekeel. Chambers stro ngly inflat ed , glo bular, rapidly increasing in size, 6-7 in number in the lastwhorl on dorsal side; cha mbers of inner who rls mor e flattened , poorly visible, fused on ventra l

FORAMIN IFE RA FROM TH E PAL EOCENE OF POLA ND 57

side. Sutures radial, stro ngly depressed on dorsal side, flush with surface, indistinct on ventra lside. Surface rough, coarsely pitted and somewhat tu bercular, especially so on the ventra l side,except for the admargina l part of it, which is almos t smooth. Aperture, a slit a t the base of thelast cha mber, near the margin of ventra l side.

Variation insignificant; it applies only to the size and development of ornam entation ;it is not well known because of the sca rcity of material.

Remarks. - Our specimens are very similar in genera l appea ra nce to those from Bundeboring (Holland) illustrated by MARIE (MS); they have similarly developed dorsal sides, whiletheir ventra l sides are difficult to compare becau se of the too schematic drawing done by MARIE.Rotorbinella papillata n. sp. is somewha t similar to R . montiana n. sp ., from which it differsma inly in lacking of an umbilicus and in being more perforated especially on ventral side.

Occurrence. - In Poland very rare, in the Pa leocene (Montian) of Pamietowo boringonl y. In Belgium it was found by the pre sent writers in the Paleocene (Montian) in Mo ns("Pu its Artesien"). Pre sent also in the Pa leocene (Montian) of Holland (Bunde).

Family SPIRILLTNIDAE REUSS, 1862

Genus PATELLINA WILLIAMSON, 1858

Patellina sp .

(PI. Ill , figs. 4, 5)

Material. - About 20 specimens badly preserved.Dimension s of two specimens (in mm):

Longest diameterShortest diameterHeight of test

F.IX/200·300·30

F. IX/210·350·300·21

Description. - Test conical , involute, dorsal side elevated, um bilical side flat or evenconcave, outline rounded. The bad state of pres ervati on does not allow all the deta ils to bedistinguished or the inn er structure of test to be an alysed, but it seems that the test con sistsof proloculum follo wed by an elon gated tube, coiled into 7 whorls. The outer margins of thewho rls are serrate-like, giving an impression of numerou s small chamberlets, which in realitydo not exist. On th e umbilical side, cro ss-like ind istinct gro ove occurs. Aperture obscuredcanno t be ob served .

Variation not known due to the poverty of material.

Remarks. - Our spec imens seem to be similar, on one hand, to those of Patellinadentata TERQu EM (1882, p. 123, PI. 12, figs. 36, 37) by their similar shape and ornamentationand, on the other hand, to those of recent species P. corrugata WILLIAMSON (1858, p. 47, PI. 3,figs. 86-89). Th ey both are too schematically illustrated to be compared with Polish specimens,but it seems tha t the main difference lies in the ornam ent at ion on the ventral side.

Occurrence. - In Pol and kn own from the Paleocene (Montian) of the Parnietowo boringonly. Probably present in the lowermost Tertiary of Holland.

58 KRYSTYNA POZARYSKA & JANINA SZCZECHURA

Genus SPIRILLINA EHRENBERG, 1843

Spirillina cf. nodifera T ERQlJEM, 1882

(PI. Ill, figs. 1-3)

1882. Spirillina nodifera TERQUEM; O. TERQUEM, Les Foraminife rcs de l'Eocenc. . ., p . 34, PI. 1, fig. 32 a-b.1948. Spirlllina nodifera T ERQUEM; F . BROTZEN. T he Swedi sh Pa lcocene. .. , p. 67, P I. 10, fig. 12 (here earlier synonymy

included) .

Material. - About 20 specime ns, rather badl y preserved .Dimensions of two specimens (in mm):

Lon gest dia meterShortest d ia meterHeight of tes t

F. IX j220·300·27

F. IX j230·220·20

0·07

Description. - Test plani spiral , evo lute, co iled in one plan, some wha t concave on bothsides, composed of a very sma ll proloculum an d a lon g, undivided , tubular seco nd chamber,forming 3-6 whorls moderat ely increasing in size as added . From as ide rectan gular, ratherthi ck. Numerous small tubercles, more regular or less, a re seen o n one side, a nd irregular,indi stinct protuberan ces on th e oppos ite side . T hey all give a false impression of the presenceof sever al , sma ll chamberlets. Surface generally rough , porous. Ape rture formed by the op enend of the tube, in most cases ob : :ured .

Variation con cern s mainly th e size a nd or namentation development , but study couldnot be mad e as nearl y a ll specimens are too badl y preserved.

Remarks. - Our specimen s, whose surface o rnamenta t ion is usually indi stinct, seem tobe most relat ed to specimens referred by T ERQUEM (1882) to Spirillina nodi/era. That is whythe specime ns from Poland a re described here as conforming to S. nodi/era T ERQuEM.

Occurrence. - In Poland not common in the Paleocene (Montian) of the Polish Lowlandsa nd Parnietowo boring. Kn o wn to be commo n in the Paleocene (Selandian) of Sweden , rarein the Eocene of France and Am erica .

Family ROTALUDA E EHRENBERG, 1839

Genus CUVILLlERINA DEBOURLE. 1955

CuviIIierina? pomeraniana n. sp.

(PI. IV, figs. 4-~)

1961. Thalmannita madrugaensis (CUSHMAN & BERMUDEZ) ; J. HOFK EJ< Foraminifera from the Cretaceo us.. ., LVI , p . 124,Tex t-fig. 4a-c (non Rotalia madrugaensis CUSHMAN & BERMUilEZ, 1947, p . 24, PI. 5, fig. 4a-c ).

1966. Thalmannita madrugaensis (CUSH MAN & BERMUDEZ) ; J . HOFKER. Maest rich tian .. ., p. 234, PI. 53, figs . 55-60 .

Holotypus: Specimen presented on PI. IV, fig. 5 (F. IX j2(;)'Paratypi: Specimen s presented on PI. IV, figs. 4, 6 (F . IX/25. 27;.Stratum typicum: Montian .Locus typicus: Pami etr-wo boring, depth 210 ·7 m (North Poland).

Derivatio nominis: La! pomeraniana - fro m Pomeran ia, name of north region of Poland.

FORAMJNIFERA BROM THE PALEOCENE OF POLAND 59

F. IX/260·300·200·15

Longest diameterShortest diameterHeight of test

Material. - A dozen or so specimens, most of them damaged.Dimensions of two specimens (in mm):

F. IX/250·200·150·15

Description. - Test very thick, somewhat plano-convex, dorsal side slightly convex,ventral side flat or even concave, outline oval, slightly lobulate, peripheral margin broadlyrounded. Chambers globular, inflated, arranged in two or more whorls, the inner ones in­distinct, 6-7 in number in the last whorl, moderately increasing in size as added. Sutures ra­dial , deeply depressed on both sides, surface extremely rough, completely covered by numerousprotuberances and knobs, forming a meander-like pattern, especially well developed and vi­sible along sutures. On both sides of test, several small knobs occur in the central part. A che­vron pattern over sutures, typical for Cuvillierina genus, can be observed but rarely. Aperture,a distinct slit on the peripheral margin under a narrow lip.

Variation insignificant; it applies mainly to the size and surface ornamentation, whichcan be more regular or less, forming meandering ribs perpendicular to the suture line, or ir­regularly arranged protuberances.

Remarks. - Our specimens do not differ from those described as Thalmannita madru­gaensis (CUSHMAN & BERMUDEz) by HOFKER (1961) from the basal sediments of the Paleocenein Belgium, neither do they differ from comparative specimens from true Montian of Belgium("Puits Artesien" in Mons), and of Holland (Bunde boring), nor from those of Paleocene ageof Germany (Krefeld region). However, all these differ from specimens described by CUSHMANand BERMUDEZ (1947) as Rotalia madrugaensis from Paleocene of Cuba and Madagascar.In the light of the above, HOFKER's specimens ought certainly to be included into our newlyerected species Cuvillierina? pomeraniana. Our specimens are also similar to those describedas Laffiteina vallensis by GAONA (1948) from the Eocene of Spain and France. They differ,however , in having somewhat different ornamentation. The last chamber of our specimensis more rounded in outline and not elongated, moreover the peripheral margin of Polishspecimens is broadly rounded in contrast with the acute periphery of Laffiteina vallensis Gaona.

Specimens of Cuvillierina? pomeraniana from Poland are only tentatively assigned toCuvillierina genus ; their somewhat asymmetrical coiling rather refers them to Cuvillierina,but on the other hand, their ornamentation is more characteristic for representatives of Thal­mannita genus.

Occurrence. - In Poland not very common in the Paleocene (Montian) in Pamietowoboring only. Known to be common in Paleocene (Montian) of Belgium ("Puits Artesien" inMons), of Holland (Bunde), Germany (Krefeld region) and probably of France (Paris Basin).Thi s species seems to be a good facies and stratigraphical index fossil for subtropical Montian.

Genus PARAROTALIA LE CALVEZ, 1949

Pararotalia tuberculifera (REUSS, 1862)(PI. VII, Figs. 1-10; Text-fig. 13)

1862. Rotalia tuberculifera REUSS; A. E. REUSS, Die Foraminiferen des Kreides.. ., p, 313, PI. 2, fig. 2.1957. Pararotalia tuberculifera (REUSS) ; J. HOFKER, Foraminifera from the Cretaceous..., XXIV, p. 32, Text-figs. 1-18.1965. Pararotalia tuberculifera (REUSS); K. POZARYSKA, Foraminifera and biostratigraphy... , p. 118, PI. 20, fig. 3a-c.

60 KRYSTYNA POZAR YSKA & JANI NA SZCZECHURA

Material. - Som e hundred specimens well preserved.Dimensions and description are given in POZARYSKA'S pap er (1965).Variation con siderabl e; it applies to the genera l sha pe, size and ornamenta tion. Outline

is regularly lobulat e up to strongly serra te, especially when cha mbers are marginally borderedby distinct, sharp spine. Umbilicus sometimes larger, sometimes less large, generally witha central plug single or divided into several sma ll, kno b-like plugs; protuberances around centralplug, if pre sent, formed by older raised parts of cha mbers (tena) . On the dorsal side, tests areeither smooth or ornamented by subradial, irr egular thickenings (see Text-fig. 13).

2

Q1 mm..-.---.3

4

F ig. 13Individual variat ion of Pararotalia tuberculifera (REUSS); Pamietowo boring , 215 m , Montian.

Remarks. - This spec ies was studied in detail s by HOFKER (1957-1963), who pr esentedits numerous variatio ns. Our specime ns belong certai nly to the same species.

Occurrence. - In Poland , not common in the Paleocene, i.e. Danian and Montian ,especially of Pami etowo boring, where it is most common . Known to be common in the Pa­leocene (Montian) of Belgium ("Puits Artesien" in M on s) and in Maastrichtian-Paleocene ofHolland (Bunde).

Ge nus ROTALIA LAMARCK, 1804

Rotalia marginata D'ORBIGNY, 1826

(PI. v, figs. 2, 5-9 ; PI. XVIII, figs. 1-4)

F. IX/290·250·20

Longest diameterSho rtest di ameterHeigh t of test

Material. - Thirty five specimens in d ifferent sta teDimensions of two specimens (in mm):

F. IX/3 30'500·470·30

1826. Rotalia marginata O'O RBIGNY; A. O'ORIlIGNY, Ann. Sci . Na t., p. 106 (272) no . 9 - nom. nudo (fide Cat. o f Fora m .,ELLIS & M ESSINA).

1850. Rotalia marginata O'O RIllGNY; A. O'O RBIGNY, Prod rome de Paleontologie... , vo l. 2, p. 407 , Figs. 1, la-b.1946. Rotalia marginat a O' O RBIGNY ; R. c. VAN BELLEN, Forarninifera fro m the M iddl e Eocene . .. , p. 63, PI. 8, figs. 11-13 .'!1946. Discorbis pseudodiscoides VA N BELLEN; R. C. VA N BELLEN, Ibid. , p. 53, PI. 6, figs. 10-15.1961. Gavelinopsis pseudcdiscoides (VAN BELLEN); J . H OFKER, Forarnin ifcra.. ., p. 124, figs. Sa- c.1966. Discorhis pseudodiscoides VAN BELLEN; J. HOFKER, Maestrich tian , D ani an .. ., p. 244 , PI. 47 , fig. l7a-c.

of preservat ion.

Description. - Test biconvex, more convex or almost conical on dorsal side, less convexon ventral side, rounded in outline, not lobulate, periphery sharply acute. Two and a half

FORAMINIFERA FROM THE PALEOCENE OF POLA ND 61

indistinct whorls on the dorsal side , the last one visible on the ventral side. Chambers stronglycompressed, 7-9 in number in the last whorl. Sutures to be seen only on the ventral side, de­pres sed and enlarged at the fairly prominent median knob. The development of median knobvaries in different specimens ; it may be connected with the walls of the chambers or in differentdegrees detached from them, moreover, it may have a small hollow on the top, in the middle.Surface smooth and glossy on the dorsal side, distinctly pitted, except the admarginal areaon the ventral side. Aperture, slit-like at the base of the last chamber on the ventral side.

Variation rather significant; it concerns above all the size of test and development of centralpart of the ventral side. The median knob may be developed in many ways; it may be highor low and only rarely is directly connected with the wall chambers, like on PI. V, fig. 6 andPI. XVIII, figs. 1--4.

Remarks. - The illustration done by D'ORBIGNY is very schematic and inaccurate, thatis why our designation is based on VAN BELLEN'S paper (1946). Our specimens represent thesame species as described by VAN BELLEN as Rotalia marginata , although our specimens aremuch smaller than those belonging to that species from Holland. Discorbis pseudodiscoidesVAN BELLEN seems to contain specimens of R. marginata , in which the median knob is connectedwith walls of chambers, as is mentioned in the description and variation of R. marginata.Thi s suggestion is confirmed when analysing variation of R. marginata specimens from the com­parative material from Montian of Holland (Bunde), (see PI. XVIII, figs. 1--4). Therefore,specimens referred by VAN BELLEN to Discorbis pseudodiscoides are here tentatively includedinto synonymy of Rotalia marginata.

Occurrence. - In Poland, common in the Paleocene (Montian) of Pamietowo boringonly. Known to be common in the Paleocene (Montian) of Holland (Bunde) and Belgium("Puits Artesien" in Mons), as well as in the Lower Eocene of France.

Rotalia saxorum D'ORBIGNY, 1850

(PI. V, figs. I, 3, 4; PI. XVIII, figs. 5-9)

1826. Rotalia saxorum O'ORBIGNY; A. O'ORBIGNY, Ann. Sci. Nat. , p. 106(272), nom. nudo (fide Cat. of Foram., EL-LIS & MESSINA).

1850. Rotalia saxorum O'ORBIGNY; A. O'ORBIGNY, Prodrome de Paleontologie , p. 407, fig. 5.1882. Rotalina perovalis TERQUEM; O. T ERQUEM, Les Foraminiferes de l'Eocene , p. 70, PI. 6, fig. 5.1946. Rotalia saxorum O'ORBIGNY; R . C. VAN BELLEN, Foramini fera from the Middle Eocene. .. , p. 64, PI. 8, figs. 17-19.1952. Rotalia perovalis (TERQUEM) ; Y. LE CALVEZ, Revision des Foraminiferes. .. , p. 49, PI. 4, figs. 47-48.1959. Rotalia saxorum O'ORBIGNY; J. HOFKER, Les Foraminifere s des cra ies. .. , p, 286, figs. 79-84.1966. Rotalia saxorum O'ORBIGNY; J. H OFKER, Maestrichtian, Danian and Paleocene..., p, 263, PI. 55, fig. 94; p, 246,

PI. 47, fig. 20a-c.1966. Rotalia perovalis (TERQUEM) ; J. HOFKER, Ibid., p, 263, PI. 55, fig. 97.

Material. - About 20 specimens in a rather poor state of preservation.Dimensions of two specimens (in mm):

Longest diameterShortest diameterHeight of test

F. IX/340·500'460·28

F. IX/350·250·20

Description. - Test plano-convex, flat on the ventral side, conical on the dorsal side,quite round in outline, not lobulate, periphery acute. Test is composed of probably more than

62 KR YSTYNA POZAR YSKA & JANI NA SZCZECHURA

two whorls, the last one containing 6-7 chambers visible on the ventra l side. The arrangementof chambers on the dorsal side is qu ite ob scured , sutures flush with surface not visible. On theventral side, sutures distinct, do not reach the margin of test, but becom e deeper and enlargednear the well pronounced umbilical plug. Surface coarsely pitted on the ventra l side aro undthe umbilicus, smooth and glossy on the dorsal side, as well as along the periphery of ventralside. Aperture, slit-like at the base of the last formed chamber, on ventral side.

Variation insignificant; it applies mainly to the general size of test and th e developmentof older raised parts of chambers (tena), which are developed in different degree. In sma ll spe­cimens, probably young ones, tena are not yet distinctly developed (see PI. V, figs. 3-4 andPI. XVIII, figs. 5-9).

Remarks. - Our specimens seem to differ only in size from those described by O'ORBIGNY(1825, emend. FORNASINI, 1906), VAN BELLEN (1946) and HOFKER (1959, 1966). Specimen il­lustrated and described by TERQuEM (1882), revised by LE CALVEZ (1952), as Rotalia perovalisfrom the Lutetian of Paris Basin, is in our opinion conspecific with R. saxorum O'ORBIGNY,falling well within the variability of thi s last one, as it was observed in th e comparati ve materialfrom Montian of Holland (Bunde) and as is shown on PI. XVII I, figs. 5-9. It seems that veryoften specimens, referred to R. perovalis (TERQu EM), in reality represent the early stages ofontogenetic development of R. saxorum O'ORB. That is why HOFKER'S illustrated specimenof R . perovalis (TERQ.) is included here into synonymy of R. saxorum O'ORB.

Occurrence. - In Poland, rare in the Paleocene (Montian) of Pami etowo boring only.Known to be common from the Maastrichtian up to Paleocene (M ontian) of Holland (Bunde),Belgium ("Puits Artesien" in Mons) and German y (Krefeld region ). Probably present in theEocene (Lutetian) of France (Pari s Basin).

Rotalia trochidiformis (LAMARCK, 1804)

(pI. v, fig. 10 ; PI. XVII, figs. 5-8)

1804. Rotalites trochidiformis LAMARCK ; J. B. LAMARCK, Ann. Mus., vol. 5, p. 184.1806. Rotalites trochidiformis LAMARCK; J. B. LAMARCK, Ibid., vol. 8, PI. 14, fig. 8a-b.?1850. Rotalia thouini O'ORBIGNY; A. O'ORBIGNY, Prod rome de Paleon tologie.. ., p. 107, (nom . nud.), see FORNASINI 1906,

Mern, R. Acad. Sci. Bologna, ser. 6, t , 3, PI. 2, fig. 7 (fide Cat. of Foram., ELLlS & MESSINA).1882. Rotalina saxorum (O'ORBIGNY); O. TERQuEM, Les Foraminiferes de l'Eocene.. ., p , 69, PI. 6, fig. 4.1946. Rotalia trochidiformis (LAMARcK); R. C. VAN BELLEN, Foraminifera from the Middl e Eocene. . ., p. 66, PI. 9, figs. 7-9.1946. Rotalia trochidiformis (LAMARCK) var. VAN BELLEN; R . C. VAN BELLEN, Ibid., p. 66, PI. 9, figs. 4-6.1946. Rotalia thouini O'ORB.; R. C. VAN BELLEN, lbid., p. 65, PI. 9, figs. 1-3.1954. Rotalia trochidiformis (LAMARCK); A. H. SMOUT, Lower Tertia ry Foraminifera. .. , p. 43, PI. I , figs. 1-6 (fide

Cat. of Foram. , ELLIS & MESSINA).1955. Rotalia trochidiformis (LAMARCK); J. HOFKER, Foraminifera from the Cretaceous.. ., XI, p. 119, Text-figs. a-c

on p. 121.1959. Rotalia trochidiformis (LAMARcK); J . HOFKER, Les Forarniniferes des craies... , p. 287, figs. 85-94.1959/60. Rotalia trochidiformis (LAMARCK); J. HOFKER, Les Foram iniferes du Cretace... , p. 376, fig. 38a-c.unpubI. Rotalia trochidiformis montensis MARIE; P. MARIE, Les Foraminiferes du Calcaire Pisolithique.. •, (MS), PI. 20,

figs. 1-5.1966. Rotalia trochidiformis (LAMARCK) ; J. HOFKER, Maestrichtian , Danian and Paleocene.. ., p. 247, PI. 47, fig. 19a-c.

Material. - A single specimen somewhat damaged.Dimensions of one specimen (in mm) :

Longest diameterShortest diameter

F. IX/2580·350·35

FORAMINIFERA FROM THE PALEOCENE OF POLAND 63

Description. - Test trochoidal, biconvex, almost round in outline, periphery distinctlyacute, not lobulate, ventral side more convex than the dorsal one. Chambers 7 in number seenon both sides, arranged in two or two and a half whorls, sutures radial, deeply depressed andslightly ramified on the ventral side, somewhat curved and flush with surface on the dorsalside. Surface smooth; coarse pits occur on the dorsal side, mainly between septal sutures,disappearing in the central part of test. Aperture not visible.

Remarks. - According to the revision of Rotalia trochidiformis (LAMARCK, 1804) bySMOUT (l954), our single specimen is very similar to specimens referred to this species. Incomparison with illustrated specimens, described as R. trochidiformis by VAN BELLEN (l946),SMOUT (l954) and HOFKER (1955, 1959, 1966), our specimen is smaller, more convex on theventral side than on the dorsal, and with not such intensively ramified sutures on the ventralside. The periphery of the Polish specimen is more acute, with umbilicus lacking. Specimensillustrated by MARIE (MS) from "Calcaire Pisolithique" of France, assigned by him to a newsubspecies, seem not to differ from the specimens known as R. trochidiformis (LAMARCK).Rotalia trochidiformis described by LAMARCK (1804) and R. thouini described by D'ORBIGNY(1826), both from the Eocene (Lutetian) of France, are known as differing each other in thepresence of a pronounced umbonal plug in specimens of R. thouini and lack of it in specimensof R. trochidiformis. As may be observed, however, on tests undoubtedly belonging to R. tro­chidiformis from the Paleocene, i.e. Montian of Holland (Bunde), the development of theircentral part of ventral side seem to be changeable to a different degrees within one species(see PI. XVII, figs. 5-8).

Occurrence. - In Poland a single specimen found in the Paleocene, i. e. Montian of theParnietowo boring only . Known to be common in the Paleocene (Montian) of Belgium ("PuitsArtesien" in Mons), Holland (Bunde), France (Paris Basin), Germany (Krefeld region), thePaleocene of India and Eocene of Syria and Iraq.

Family ELPHIDIIDAE GALLOWAY, 1933

Genus ELPIDDIUM DE MONTFORT, 1808

Elphidium cf. lamarcki (D'ORBIGNY, 1826)

(PI. IX, figs. 1-5)

1826. Nonionina lamarckii O'ORBIGNY; A. O'ORBIGNY, Ann. Sci. Nat ., p. 128 (294) nom. nudo (fide Cat. of Foram.,ELLIS & MESSINA).

1904. Nonionina 'lamarck ii O'ORBIGNY; C. R. FORNASINI, Mem. R. Acad. Sci . Bologna, PI. 3, fig. I .

Material. - About 40 specimens, in most cases well preserved.Dimensions of specimens (in mm):

Longest diameterShortest diameterHeight of test

F. IX/530·220·20

F. IX/550·270·220·12

Description. - Test planispiral, involute, biconvex, nearly round in outline, peripheryabruptly acute, slightly keeled. Chambers 12-14 in number in the last whorl, separated bystrongly curved, slightly limbate, depressed sutures, on which there are more distinct or less

64 K.RYSTYN A POZAR YSK.A & JAN INA SZCZ ECHURA

distinct, small openings. Umbilical region depressed, covered by numerous, regular , small pa­pillae, extending on the periphery at the base of the last formed chamber. Aperture developedas a row of pore s along the base of the last chamber.

Variation insignificant ; it applies mainly to the size, number of chambers and developmentof sutural pore s, as well as to the papillae within the umbilical region.

Remarks. - Our specimens are similar to figured specimen described by d'ORBlGNY(1826) as Nonionina lamarckii, however, this last one is illustrated very schematically. Oursare more compressed laterally having less chambers, and for that reason they are referred asconformed to Elphidium lamarcki. Our specimens resemble somewhat, in their general appear­ance, specimens described by VAN BELLEN (1946, p. 42, PI: 3, figs. 22-25) as Nonion geleenense.They differ, however, in having distinctly depressed, fairly tuberculated umbilical region .

Occurrence. - In Poland, common in the Paleocene (Montian) of Polish Lowlandsand Pamietowo boring. Present in the Paleocene (Montian) of Belgium ("Puits Artesien" inMon s) and Holland (Bunde) and in the Lower Tert iary sediments of France.

Genu s ELPffiDIELLA CUSHMAN, 1936

Elphidiella prima (TEN DAM, 1944)

(PI. IX, figs. 16-19)

1944. Elphidium primum TEN D AM ; A. TEN DAM, Die stratigraphische . .. , p . 109, PI. 3, fig. 15.1946. Nonion multisuturatum VAN BELLEN; R. C. VAN BELLEN, Foraminifera fro m the Middle Eocene... , p. 44, PI. 4 ,

figs. 4-6.1948. Elphidiella prima (TEN DAM) ; F . BROTZEN, T he Swedish Paleocene... , p. 70, PI. 8, fig. 2.1965. Elphidiella prima (TEN D AM); J. INDANS, Mikrofaunisti sches Normalprofil. .. , p. 12, PI. 1, fig. 6.

Material. - About 50 specimens well preserved.Dimensions of two specimens (in mm):

Longest diamet erShortest diameterH eight of test

F . IX/610·300·25

F. IX/630·600·550·36

Description. - Test plani spiral, involute, convex on both sides, round and indistinctlylobulate in outline, periphery rounded or slightly angled . Chambers 12 in number in the lastwhorl , moderately increasing in size as added. A distinct elevated umbo is developed in the um­bilical region. Sutures slightly depressed between the last chambers and flush with surfacebetween the chambers in the early portion of test , with two rows of pore s. Aperture invisible,surface smooth, glossy.

Variation insignificant; in small specimens tiny papillae like tubercles occur at the innermargin of the last chamber, while sutural openings and umbo are indistinct.

Remarks. - Specimens described as Elphidiella prima from Poland differ from thosedescribed as Elphidium primum by TEN DAM (1944), or as Elphidiella prima by VAN BELLEN (1946)and BROTZEN (1948) but only in having generally somewhat less chambers in last whorl.

Occurrence. - In Poland , common in the Paleocene (Montian) of Polish Lowlandsand Pamietowo boring. Known to be common in the Paleocene , i, e. Danian and Selandianof Sweden, Montian of Holland (Bunde) , France and Germany (Krefeld region) .

FORAMINI FERA FROM TH E PAL EOCENE OF POLAND 65

Family GLOBOROTALIIDAE CUSHMAN, 1927

Genus GLOBOROTALIA CUSHMAN, 1927

Globorotalia globigeriniformis VAN BELLEN, 1946

(PI. VII , figs. 11-19)

F. IX /660·200·17

Longest diamete rSh ortest diameter

Height of test

120 specimens well pre served.specimens (in mm):

F. IXf710·350 ·30

0·16

1946. Globorotalia glo bigeriniformis VAN BEI.LEN; R. C. VAN BEI.LEN, Foraminifera from the Middle Eocene . . .• p . 71.

PI. 10, figs. 10-12 .?196 1. Pararotalia globigeriniformis (VAN BELLEN); J. HOFK ER, Fora m inifera from the Cret aceous.. ., LI V, p. 86 , fig. 2a-c.?1962. Pararotalia globigerinifo rmis (VAN BELLEN); J. HOFKER, Correlation.. ., p . 1082, Text-fig. 268.

1966. Pararotalia globigeriniformis (VAN BFLLEN); J . HOFKER. Maest rich tian, Dani an a nd Pale ocene... , p. 233 , PI. 44 ,

fig. 123 a-c.

Material. - AboutDimen sion s of two

Description. - Test piano-convex, thick, rounded in outline, strongly lobulate in adultor angled in juvenile forms, periphery broadly rounded. The ventral side moderately convex,dorsal side nearly flat. Chambers globular, 6-7 in number in the last whorl , arranged in oneor two whorls; only the outer whorl to be seen on the ventral side . Sutures depressed on bothsides, more on the ventral one, especially near the umbilicus, radial or slightly oblique betweenthe last chambers. On the dorsal side there occur a rib-like projections in juvenile forms, orknob-like projections in the adult forms. Umbilicus generally filled by a single or several knobs,bordered by the older raised parts of chambers (tena). Surface rough, especially in the centralpart of the ventral side , distinctly porous. Aperture as a slit at the base of the last formedchamber, on the ventral side.

Variation considerable; it applies to the size, general shape and ornamentation. Theoutline varies from the sharply angled up to the regularly, gently lobulate. Chambers notglobular, compressed in the juvenile forms up to prominently globular in the adult, ornamen­tation more intensively developed in the adult form s, who se sur face is also more rough (seePI. VII, figs. 11-18).

Remarks. - General appearance of specimens, described as Globorotalia globigerini­formis from the Montian of Poland, seems to be very similar to the illustrated forms describedby VAN BELLEN (1946) as a G. globigeriniformis from the Eocene (recte Montian) of Holland.Our specimens do not differ from those described by HOFKER in 1966, who revised VAN BELLEN'Soriginal collection. However, Polish specimens are markedly smaller and they have less chambersin the last whorl. Moreover, generally they have a plug in the central part of the ventral side .Specimens, described by HOFKER (1961, 1962) as Pararotalia globigeriniformis, from the layerslying just above the "Md" horizon in Holland, are her e only tentatively included into synonymyof Globorotalia globigeriniformis. In comparison with specimens described by VAN BELLENand with those occurring in Poland, they are too sharply angled in the outline and they differin the development of umbilicus.

Occurrence. - In Poland, very common in the Paleocene (Montian) of Polish Lowlands(Sochaczew boring) and Parnietowo boring. Known as common in the Paleocene (Montian)of Belgium ("Puits Artesien" in Mons), Holland (Bunde), and Germany (Krefeld region).Pa laeontolog ia Polonica No. 20 5

66 KR YSTYNA POZAR YSKA & JANIN A SZCZECH URA

Globorotalia praepseudomenardii HOFKER, 1961

(PI. VI, figs. 7-10)

1961. G/oborotalia praepseudomenardii HOFKER; J . HOFKER, Foraminifera from the Cretaceous , XIV, p. 85, fig. 1a-d.1961. G/oborotalia praepseudomenardii HOFKER; J. HOFKER, Les Foraminiferes planctoniques , p. 56.1966. G/oborotalia praepseudomena rdii HOFKER ; J. HOFKER, Maestrichtian, Danian and Paleocene. .. . p. 233, PI. 43,

fig. 105; PI. 46, fig. 140.

Material. - About 50 specimens, generally somewhat damaged.Dimensions of two specimens (in mm):

Longest diameterShortest diameterHeight of test

F. IX/830·300·230·10

F. IX/810·230'18

Description. - Test compressed, flatly biconvex, trochoidal, slightly ovate in outline,periphery lobulate, bluntly acute, keeled. Dorsal side, with all chambers visible in last whorl,more convex than ventral side. Chambers gently inflated, especially on the ventral side in theouter whorl, arranged in two whorls, 8 in number in the last whorl , rapidly increasing in sizeas added. Sutures radial on the ventral side, curved or even sickled on dorsal, more depressedon ventral side. Umbilicus distinctly developed, generally filled up with a single or severalknobs, formed by older raised parts of chambers (tena). Surface slightly rough, wall porous.Aperture a slit-like on the ventral side, at the base of the last formed chamber.

Variation insignificant; it applies to the size, general shape and ornamentation. The smallertests are less lobulate in outline, with less pronounced particular chambers, moreover theyhave a singular umbilical plug. Dorsal side is more porous or less, or even covered by papillae­like ornamentation.

Remarks. - In comparison with the illustrated holotype of Globorotalia praepseudo­menardii HOFK.ER (1961), our specimens differ only in having the umbilical plug and in beingnot so high. Some specimens represented by KAASSCHIETER as Rotalia audouini D'ORB. (1961,p. 241, PI. 16, figs. 8-10), seem to be very similar to specimens assigned by the present writersto G. praepseudomenardii HOFKER. However, our specimens are never spinulose, i.e. neverornamented with spines along the periphery and they have a very distinct furrow around theumbilical plug.

Globorotalia praepseudomenardii seems to be not related with G. pseudomenardii as maybe suggested by its name; thi s last one described by BOLLI (1957), has different general appear­ance, and particularly differently arranged chambers.

Occurrence. - In Poland, common in the Paleocene rMontian) of the Pamietowo boringonly. Known to be common in the Montian of Belgium ("Puits Artesien" in Mons), Holland(Bunde), France, and of Germany (Krefeld region). This species seems to be a good index fossilfor beds of Montian age.

Globorotalia sp.

(PI. VI, figs. 1-4)

1946. Rotalia? marginata D'ORBIGNY; R. C. VAN BELLEN, Foraminifera from the Middle Eocene.. ., p. 63, PI. 8. figs. 8-10(non Rotalia marginata D'ORDIGNY, 1850, p. 407, type figure. see FORNASINI. 1906, PI. 2, fig. 1).

FORAMINI FERA FROM THE PAL EOCENE OF POLAND 67

Longest diameterShortest diameterHeight of test

Material. - Nine specimens well preserved.Dimension s of two specimens (in mm):

F. IX/850·250·200'10

F. IX/880-400·32

Description. - Test piano-convex, slightly convex on the dorsal side, flat or even concaveon ventral side, nearly round and gently lobulate in outline. Periphery bluntly acute, somewhatkeeled . Nearly two whorls visible on the dorsal side, out of which only last one is seen on theventral side. Chambers distinct , slightly inflated on both sides, especially the last ones , 6 innumber in the last whorl. Sutures somewhat cur ved, depressed on ventral side, nearly flushwith surface on dorsal side. A shallow small umbilicus, usually filled by a low plug , occurs onventral side. Surface smooth, on ventra l side, except the admarginal periphery, distinctly pitted.Aperture, a slit-like between the periphery and the umbilicus, generally poorly visible.

Variation not kno wn due to the scarcity of material. The studied specimens differ onlyin size and development of the umbilical area .

Remarks. - Our specimens are the most similar to those illustrated and described byVAN BELLEN (1946) as Rotalia ? marginata D'ORB. from -the Eocene (recte Montian) of Holland.Polish specimens differ from them only in being less conical on the dorsal side. They differ,however , con spicuously from specimens referred to R. marginata - species erected by D'OR­BIGNY.

Occurrence. - In Poland, rare in the Paleocene (Montian) of the Pamietowo boringonly. Found by the present authors in the Paleocene (Montian) of Holland (Bun de) and Belgium("Puits Artesien" in Mons). Present in Calcaire pisolithique (Montian) of France.

Genus GLOBOROTALITES BROTZEN, 1942

Globorotalites granulatus n. sp.

(PI. VIII , figs. 1-7)

1962. Globorotalites ? n. sp. ; M. E. SCHMID, Die Foraminiferenfauna... , p. 350, PI. 6, fig. 8a-c.

Holotypus: Specimen presented on PI. VIII, fig. 7 (F. IX/80).Paratypi: Specimens presented on PI. VIII , figs. I, 6 (F. IXj74, 79) ; Pamie towo boring, 274·5 m.Stratum typicum: Montian .Locus typicus: Sochaczew boring, depth 194 m (Central Poland).Derivatio nominis: Lat. granulatus - from the granulation, characteristic for the umbilical part of test.

Material. - About 80 specimens, some of them damaged.Dimension s of 3 specimens (in mm):

Longest diameterShortest diameterHeight of test

F. IX/800·300·280·18

F. IX /740·220'17

F. IX/190·350·250·22

Description. - Test trochospiral, usually high , outline nearly rounded, gently lobulate,periphery acute, slightly keeled. Dorsal side flat or even convex, ventral side distinctlyconvex. Chambers angular, inflated on the ventral side , arranged in 2-3 whorls; five chambers

5*

68 KR YSTYNA POZARYSK A & JA N1NA SZCZEC HU RA

of the last whorl increa se rapidly in size as added. Sutures slight ly depressed ventra lly, flushwith surfa ce dorsall y, genera lly curved, even sick led on th e vent ra l side . U mbilicus more deepor less, filled by sma ll papillae, or o pe ned a nd then bordered by well de veloped inner raisedparts of chambers (tena ). Surface smooth, glistening. Aperture, an enlarged slit on the ventralside, at the base of last cha m ber. near th e umbilicus, pa rtly obscured by umbilical papillae.

Variation rath er significa nt, concerning both th e size, co iling and genera l appearanceof test. Within the studied specime ns there a re forms with quite flat or somewha t convex dorsalside, with more elevated or less eleva ted cha mbe rs on ventra l side and with large, deep, welldeveloped umbilicus or with an indistinct umbilicu s, and then without distin gui shed tena.This last one depends probably on the stage development of test. Umbilical papillae are alsodifferently scattered inside th e umbilicus.

Remarks. - Our specime ns a re co mpa ra ble to Globorotalia angulata (WHITE), on onehand, and with G. acuta TOULMIN, on the other. From the first th ey differ in havin g a more acutekee led periphery, less depressed and not radial sutures and in having fairly sm ooth surface.In contrast to speci mens referred to G. acuta, our spec ime ns have not so thickened keel andsutures on the dor sal side and have chamber to ps (tena) glistening, without any ornamentation.Contrarily to two a bove mentioned spec ies, our speci me ns have rather characteris tic papillaeinsid e th e umbilical dep ression , which are lacking in other spec ies of Globorotalites genus.Specimen s of G. granulatus, especia lly those you ng ones , resemble figured specimen of Glo­borotalites ? n. sp ., described by SCHMID (1962) fr om th e Danian a of Austria . Both forms, fromPoland and Austri a , a re very similar in general appea ra nce , but as results of descr iption andillu stration of Globorotalites ? n . sp . specime ns, they lack papillae o r a sort of tubercles insidetheir umbilicu s ; th ere is, however, no difference between these probably conspecific forms whentake under consid eration SCHMID'S origina l co llectio n. Close to Globororalia ko chi PUPERfrom Upper Eocene .

Occurrence. - In Poland, common in the Paleocene (Montian) of Pam iet owo and PolishLowlands. Present in D ani an 3 of Austri a . This spec ies seems to be a good index fossil forMontian Beds.

Family GLOBlGERINIDAE CA RPENTER, P ARK ER & lONES, 1862

G enus GLOBIGERINA D'ORB1GNY, 1826

Globigerina aff. compressa P LUMMER, 1926

(Text-fig. 14)

1926. Globigerina compressa PLUMI<. ER; H . J . PLUMMER, Foraminifera of the Midway... , p. 135, PI. 8, fig. 11.1953 . Globigerina compressa PLUMMER var. compressa PLUMMER; N . N . SUBBOTINA, G lobiger inidy ..., p. 56, PI. 2, figs. 2-6.1955. Globorota lia compressa PLUMMER ; J . CUVILLlER, F. DALBlEZ & C. GLlNTZBOECK EL, Et udes micropa leonto logiques. . .,

PI. 1, fig. 3a- c.1957. Globoro talia compressa PLUMMER ; H. M. BOLLI, The genera Globigeri na.. ., p. 77, PI. 20, figs. 21-23.1957. Globigerina compressa PLUMMER; J. c. TROELSEN, Some planclo nic Foraminifera .. ., p. 129, PI. 30, fig. Su-e.1960. Globorotalia compressa (PLUMMER); R . K. OLSSON, Foraminifera of latest Cretaceous. .. , p. 45, PI. 8, figs. 20-22.1961. Globoro ta lia comp ressa (PLUMMER); P . J . BERMUDEZ, Contrib ucio aI estudio .. ., p. 1285, PI. 14, fig. 8a-b.

3 The Danian age of " Bruderndorfer Feinsandes" from Austria is called in question here by the present writers.

FO RA MIN IF ERA FRO M THE PALEOCEN E OF PO LA ND 69

Lon gest diam eterShortest diameterHeight of test

1962. Globorotalia ( Turborotalia) comp ressa (PLUMMER); W. A. BERGGREi':, So me plancto nic. .. , p, 94, PI. 14, fig. 5a -c ;

Text-fig. 13 (1-6).1962. Globigerina compressa PLUMMER ; M. E. SCHMID, Die Foram iniferenfa una des Bruderndorfe r. . " p. 347, PI. 6,

fig. 6 a-c.

1965. Globigerina sp.; K . POZARYSKA, Foraminifera a nd biostrat igraphy , p. 126, PI. 22, fig. 1a-c .1966. Globorotalia compressa (PLUMMER); Z. R. EL-N AGGAR, Strat igraph y , p. 203, PI. 17, figs. 1-3.no n 1966. Globigerina compressa PLUMMER; .I. HOFKER, Maestr ich tian , D anian and Paleoccne... , p . 231, PI. 43, fig. 94.

Material. - A do zen or so specimens , often dam aged .Dimensions of an ave rage spec imen (in mm):

F. VI/ I270,22

0·190·\2

Description. - Test small, very low , trochospiral , pian o-convex, somewhat compressed ,outline lobulat e, periphery br oadly rounded or ac ute. Dorsal side flat. ventra l co nvex . Cha mbers

:1Fig. 14

Indi vidua l variation of Globigerina aff. compressa PLUMMER ; Parnietowo boring, 241 m, Montian .

inflated, sometimes globular, genera lly 5 in number in th e last whorl , rapidly increasing in sizeas added. Sutures di stinctly depressed. U mbilica l depression narrow and deep . Aperture, a lowarched o pening, extending from th e umbilicu s toward s th e periphery, bordered by a distinctlip . Wall smooth, very finely perforat ed .

Globigerina daubjergensis BRONNIMANN, 1953

1953. Globigerina daubjergensis BR6NN1MANN; P. BRONN1MANN, No te on pla ncionic. .. , p. 340, Text-fig. 1.1962. Globigerina daubjergensis BRONNIMANN; M. E. SCHMID , Die Fo ram inife renfauna.. ., p. 346, PI. 6, fig. 4a-c.

1965 . Globigerina daubjergensis BRONNIMANN; K. POZARYSKA, Fo raminifera and biostrat igraphy. .. , p. 121, PI. 23, fig. 4a-c(here additional synonymy included) ,

1966. Globigerina daubjergensis BRONN1MANN; Z. R. EL-NAGGAR, Stra tigraphy. .. , p, 161, PI. 15, fig. 3u-c .

70 KRYSTYNA POZARYSKA & JANINA SZCZECHURA

Material. - A few specimens well preserved .D imen sions, description, va ria tion and remarks are given in POZARYSKA'S paper (1965).

Occurrence. - In Poland , in the Paleocene, i. e. Danian and M o nt ian , where it is lesscommon, of Pamietowo boring and Polish Lowla nd s. This species is an ind ex fossil fo r theDanian bed s a ll over the wo rld .

Globigerina kozlowskii BROTZEN & POZARYSKA, 196 1

(Text-fig. 15)

1916. G/obigerina ko z/o.....skii BROTZEN & POZARYSKA; F. BROTZEN & K. POZARYSKA, Foraminiferes du Paleocene .. ..p. 162, PI. I , figs. 1-14; PI. 2, figs. 1-17.

1962. G/obigerina koz/o.....ski i BROTZEN & POZARYSKA; J. HOFK ER . Foraminifera from the Cretaceous... , LXI, p. 129.Text-fig. on p, 130.

1965. G/obigerina ko z/o.....sk ii BROTZEN & POZARYSKA ; K . POZARYSKA, Foraminifera and biostratig raph y..., p. 122.PI. 23, fig. I a-c.

1966. Globigerina koz lo skii BROTZEN & POZARYSKA; Z. R . EL-NAGGAR, Stratigrap hy.. ., p. 168, PI. 15, figs. I a-c, 2.1966. Globigerina koz lo sk ii BROTZEN& POZARYSKA; J. HOFKER, Macstrichtia n, Danian and Paleocene.. .. p. 231, PI. 46,

fig. 139.

Fig. 15Individual variation of Globigerina ko zlowsk ii BROTZEN & POZARYSKA; Pamietowo boring, 243 m, Montian.

Material. - One hundred or so specime ns well preserved .Dimensions, description , a deta iled va riat ion a nd remarks are given in th e paper by

BROTZEN & POZARYSKA (1961) a nd POZARYSKA (1965); add itiona l exa mples of infraspecificvariation are given above (Text- fig. 15).

Occurrence. - In Poland co mmon in the Pale ocen e, i. e. Montian of P amietowo boringand Polish Lowlands. Present in the Paleocen e of Holland (Limburg region and Bunde boring).Noted in Egypt by EL-NAGGAR (1966) as ea rly as in the uppermost D anian up to the LowerPaleocene.

Globigerina pseudobulloides PLUMMER, 1926

1926. G/obigerina pseudobulloides PLUMMER ; H. J. PW MMER, Foraminifera of the Midway.. ., p. 133, PI. 8, fig. 9.1962. Globigerina pseudobulloides PW MMER; M. E. SCHMID. D ie Foraminiferenfauna .. ., p. 348, PI. 6, figs. I a-c, 2.1965. G/obigerina pseudobulloides PLUMMER; K. POZARYSKA, Foraminifera and biostratigraphy.... p. 123. PI. 23, figs. 2-3.1966. Globorotalia pseudobulloides (PLUMMER); Z. R. EL-NAGGAR. Stratigraphy.. ., p, 224, PI. 18, fig. 3a-c.

FORAMINIFERA FROM THE PALEOCENE OF POLAND 71

Material. - About 50 specimens, rather well preserved.Dimension s, description, variation and remarks are given in POZARYSKA'S paper (1965).

Occurrence. - In Poland common in the Paleocene, i.e. in Danian an d Montian of Pol ishLo wland s an d Pamietowo boring. Common from upperm ost Cretaceou s up to the LowerTertiary all over the world.

Globigerina (Subbotina) triloculinoides PLUMMER, 1926

1926. Globigerina triloculinoides PLUMMER ; H. J . PLUMMER. Foramini fera of the Midway... , p. 134, PI. 8, fig. 10.1965. Globigerina (Subbotina) triloculinoides PLUMMER ; K. POZARYSKA, Fo rami nifera and biostra tigraph y. .. , p, 124.

PI. 22, fig. 2a-c (here additional synonymy included) .1966. Globigerina triloculinoides PLUMMER; Z. R. EL-NAGGAR. Strati graphy.. .. p. 178, PI. 15, fig. Ta-c.

Material. - About 30 specimens well preserved .Dimension s, description , variation and remark s are given in POZARYSKA'S paper (1965)

and a detailed discussion in EL-NAGGAR'S pap er (1966).

Occurrence. - In Poland not so common in the Paleocene, i. e. Danian and Montian ,where it is more comm on tha n in Danian , of Polish Lowlands and Parnietowo boring. Knownto be common from the Dani an up to the Eocene from some countries in Europe and America.Restr icted to the Paleocen e in Egypt (Esna-Idfu region), where is considered as on e of the bestindex fossils for the Paleocene.

Globigerina trivialis SUBBOTINA, 1953

1953. Globigerina trivialis SUIIBOTINA; N. N. SUBIIOTlNA, Globigerinidy.. ., p. 64, PI. 4. figs. 4-6, non figs. 7-8.1965. Globigerina trivialis SUBBOTlNA; K. POZARYSKA, Foraminifcra and biostratigraphy.. ., p. 125, PI. 22, fig. 3a-c

(here additional synonymy included) .

Material. - Several specimens well preserved.Dimensions, description , variation and remarks a re given in POZARYSKA'S paper (1965).

Occurrence. - In Poland rare in the Paleocene, i.e. Danian and Montian of PolishLowlands and Pami etowo boring. Kn own from the uppermost Maastri chti an up to the Danianin U. S. S. R. (Russian Platform ).

Globigerina varianta SUBBOTINA, 1953

1953. Globigerina varianta SUBIIOTlNA; N. N. SUBBOTINA, Globigerinidy... . p. 63, PI. 3, figs. 5, 10-12; PI. 4, figs. 1-3;PI. 15, figs. 1-3.

1965. Globigerina varianta SUBBOTlNA; K . POZARYSKA, Foramin ifcra and biostrat igrap hy. .. , p. 125, PI. 23. fig. 5a-c(here additional synonymy included) .

Material. - Several specimens well preserved.Dimension s, description , variation and remarks are given in POZARYSKA'S paper (1965).

Occurrence. - In Poland rare in the Pal eocene, i. e. Dan ian and Montian of Polish Low-lands and Pamietowo boring. Known to be common in Europe in the Lower Tertiary sedimentsup to the Eocene.

72 KRYSTYNA POZA RYSKA & JANINA SZCZECHU RA

Family EPONIDI DAE HOFKER, 1957

Genus EP ONIDES DE MONTFORT, 1808

Eponides toulmini BROTZEN, 1948

(PI. X V, figs. 1-4)

1946. Epouides gratus (Rsuss) : R. C. VAN BELLEN, Foraminifcra from the Middle Eocenc. . ., p. 57, PI. 7, figs. 4-9 (nonRotalia grata REUSS, 1865, p. 47, PI. 4, fig. 17).

1948. Eponides toulmini BROTZEN; F. BROTZEN, The Swedish Paleocene... , p. 178, PI. 10, fig. 16.1959. A labamina bigibbera LE CALVEZ; Y. LE CALVEZ, Etude de quel qu es Forarniniferes..., p, 94, PI. I , figs. 5- -7.1960 . Eponides veslensis ROUVILLOIS ; A. ROUVILLOIS, Le Th anetien . .. , p. 68, PI. 2, fig. 29a-b.1961. Eponides toulmini BROTZEN; J. P. H. KAASSCHI ETER, Foramini fcra or the Eocene , p. 211, PI. 12, fig. La-c,1965. Eponides toulmin i BROTZEN; K. POZARYSK A, Foraminifera and biostratigrap hy , p. 110, PI. 17, fig. 4a-c.1965. Eponides gratu s (REUSS) ; J . INoANs, Mikrofaunistisches Norma lprofl l.. ., p. 13, PI. 1, fig. 2.1966. Eponides toulmini BROTZEN: J. HOFK ER, Maestrichtian , Dani an and Paleocene... , p, 261, PI. 54, figs. 83-84.

Material. - Some hundred specimens well preserved.Dimensions, description and remarks are given in POZARYSKA'S paper (1965) .

Variation rather significant ; there are forms coiled left as well as right , with peripheryacute to different degrees and round or di stinctly lobulat e outline. Te sts d iffer in height andnumber of chambers. J uvenile specimens have less coils th an adult ones and th ey are not so high(see PI. XV , figs. 1--4).

Remarks. - According to rem arks in POZARYSKA'S paper (1965) , it mu st be added hereth at the specimens, described by ROUVILLOIS (1960) as Eponides veslensis , and specime n fig uredby LE CALVEZ (1959), described as Alabaniina bigibbera n. sp., rep resent undoubtedl y Eponidestoulmini BRoTz EN.

Occurrence. - In Poland , very common in th e uppermost Cretaceous and the Paleocenc,i. e. Da nian and Montian o f Poli sh Lowl ands a nd Parnietowo boring. Known as commonfrom the Upper Maastrichtian up to the Pale ocene (Montian) of H oll and , Belgium, France,Germa ny a nd U. S. S. R ., from the Palcocene, i. e. Thanetian of France (Paris Basin) and Englan d.

Gen us NE OEP ONIDES REISS, 1960

Neoeponides cf. schrei bers i (D'ORBlGNY, 1846)

(PI. x v, fig. 5)

1846. Rotalina schreibersii D'ORBIGNY; A. O'ORBIGNY, Foraminiferes fossilcs. .. , p. 154, PI. 8, figs. 4-6.1960. Neoeponides schreibersi (D'ORBIGNY); Z . R EISS, Structure of the so-called Eponides.. ., p. 17.1961. Eponides schreibersi (O'ORBIGNY); J. P. H. KAASSCHIETER, Fo rami nifera fro m the Eocene... , p. 210, PI. 11 , figs. 14,

15a-c.

M aterial. - Two specimens well preser ved.D imensions of one specimen (in mm):

Longest diameterShortest diamet erHeight of test

F . IX/930·230·200' 11

FORAMINIFERA FROM TH E PALEOCEN E OF POLAND 73

Description. - Test small, trochospiral, piano-convex, high on dorsal side, nearly flaton ventral side, outline lobulate, rounded, periphery angled and carinate, about 3 whorlspoorly visible on dorsal side; ventral side involute, with triangular chambers slightly inflated,9-10 in number in the last whorl. Sutures indistinct in the early portion of whorl, slightly de­pressed between last chambers; spiral suture fairly thickened , septal sutures oblique, flush withsurface on dorsal side, radial on ventral side. A small, distinct umbilical depre ssion, surroundedby weakly developed secondary thickenings of older raised parts of chambers (tena) nearumbilical margin , forming there a star-like pattern. Surface smooth, glassy, finely porous.Aperture poorly visible as an interiomarginal slit-like arch, extending from the periphery toumbilicus.

Variation not known due to the scarcity of material.Remarks. - Our specimens seem to be the most similar to the figured specimen of

Rota/ilia schreibersi (recte Neoeponides schreibersi), species described by D'ORBIGNY (1846)from Miocene of Vienna Basin. Polish specimens in comparison with that from Austria differ,however, in being smaller, less biconvex, and in having thickened spiral suture. Tena are not sodistinctly developed. In the Paleocene, i.e. Montian of Holland (Bunde and Beatrix borings)and in the Eocene of Belgium (see KAASSCHI ETER, 1961) there are specimen s more similar tothat from the Miocene of Vienna Basin, so it is quite possible that in Poli sh Montian ther e occurthe most primiti ve form s, repre senting a starting line of the above mentioned species. A detailedrevision of thi s species is given by REISS (I960), after whom systematic position of the abovediscussed species is maint ained here.

Occurrence. - In Poland, two specimens in the Paleocene , i. e. Montian of the Pamieiowoboring only. Present in the Paleocene (Montian) of Holland (Bunde and Beatrix borings)according to present writers' own observation. Known in the Eocene of Belgium and Mioceneof Austria (Vienna Basin), Israel and Morocco.

Family CIBICIDIDAE CUSHMAN, 1927

Genu s CIBICIDES DE MONTFORT, 1808

Cibicides asteroides n. sp.

(PI. XI, figs. 3-7)

?1961. Cibicides sp. cr. mauricensis HOWE& ROBERTS; J. P. H. KAASSCHIETER. Foraminifera of the Eocenc..., p. 223,PI. 14, fig. 8.

19(,5. Cibicides sp.; K. POZARYSKA, Foraminifera and biostratigraphy..., p. 138, PI. 28, fig. 3a-£' .

Holotypu s: Specimen presented on PI. XI , fig. 7 (F. IX/98).Paratypi : Specimens presented on PI. XI , figs. 5, 6 (F. IX/96, 97).S tratum typicum : Montian.L OCIIS typicus: Pamietowo boring, depth 21 5 m (North Poland).Derivatio nominis: Lat. asteroides = star-like; after a star-like pattern, developed in the central part of the

ventral side.

Longest diameterShortest diameterHeight of test

cases well preserved.Material. - About 50 specimens, in mostDimensions of 3 specimens (in mm):

F. IX/960·250·22

F. IX/970·420·35

F. IX/980·350·250·15

74 KRYSTYNA POZARYSKA & JANINA SZCZECHURA

Description. - Test piano-convex, almost round and gently lobulate in outline, peripheralmargin angular. Dorsal side, nearly flat, bears a small plug, surrounded by a more distinct orless distinct furrow, penetrating somewhat into the inner margin of chambers and formingthere a star-like pattern. The ventral side st ro ngly convex, with a slightly depressed plug.Chambers 10-12 in number in the last whorl, the only one seen on dorsal side, increasinggradually in size as added. Sutures distinct, rather depressed and radial , only very slightly curvedon the ventral side, elevated, st rongly limbate and curved on dorsal side. Surface perforated,more coarsely on the dorsal side, finely on ventral one. Aperture slit-like, marginal, extendinghowever onto the ventral side, as well as on the dorsal one, sometimes bordered by a narrow lip.

Variation insignificant. It applies only to the size and the development of the umbonalregion. Adult forms are sometimes somewhat uncoiled.

Remarks. - Specimens referred to Cibicides asteroides n. sp . from the Montian of Polandseem to be similar in general appearance to specimens, described by KAASSCHIETER (1961)from the Eocene of Belgium, included by him tentatively to Cibicides mauricensis Hows & Ro­BERTS. Belgian specimens seem to differ only in having not thickened, depressed sutures on thedorsal side.

Occurrence. - In Poland common in the Paleocene, i.e. Montian of Polish Lowlandsand Pamietowo boring. It ha s been observed in samples from the Lo wer Eocene of France,and probably in the Eocene of Belgium.

Cibicides aurouzae ROUVILLOIS, 1960

(PI. x, figs. 8-12)

1946. Cibicides choctavensis CuSHMAN& McGLAMARYvar. ornata VANBELLEN; R. C. VANBELLEN, Foramini fera fromthe Middle Eoccne... , p. 79, PI. 12, figs. 7-9.

1960. Cibicides ( Cibicidina) aurouzae ROUVILLOlS; A. ROUVILLOlS, Le Th anetien .. ., p. 76, PI. 4, fig. 60 a-c.1965. Cibicides ornatus (VANBELLEN) emend. POZARYSKA; K. POZARYSK A, Foraminifera and biostratigraphy..., p. 133,

PI. 25, fig. I a-c.

Material. - Some hundred specimens well pre served.Dimensions, description and remarks are given in POZARYSKA'S paper (1965).

Variation considerable; it applies to the size and general appearance, as well as to thestructura l details of test. The most variable is the development of umbilical region, wherethere is a star-like regular hollow in small, probably young specimens, and a large, irregularopening, often bordered by thickenings of older rai sed parts of chambers (tena) in large spec­imen s. The umbilical depression is very often filled up by numerous tubercles.

Remarks. - According to remarks given in POZARYSKA'S paper (1965), it ought to bementioned here that specimens, described by ROUVILLOIS (1960) a s Cibicides aurouzae from theThanetian of France, in reality do not differ either from those described by VAN BELLEN asC. choctavensis ornata and later by POZARYSKA as C. ornatus, or from those referred toC. aurouzae by the present writers.

Occurrence. - In Poland very common in the Paleocene, i.e. Montian of the PolishLowlands and Parnietowo boring. Common in the Paleocene, i.e. Montian of Holland (Bunde)and Belgium ("Puits Artesien" in Mons), in the Paleocene, i, e. Thanetian of France (ParisBasin) , as well as in Lower Eocene of Pari s Basin (writers' own observation).

FORAMINIFERA FROM THE PAL EOCEN E OF POLAND

Cibicides carinatus (TERQUEM, 1882)

(PI. XI , fig. I ; Te xt-fig. 16)

75

Longest diameterShortest diam ete rHe ight o f tes t

1882. Truncatulina carinata TERQUEM ; O. T ERQUEM, Les Foraminiferes de l' Eocene... , p. 94, PI. 10, figs. 1-2.1944. Cibicides orbicularis (T ERQUEM); A. TEN D AM, Die strat igra phisehe .. ., p. 134, PI. 5, fig. lOa-c.1949. Cibicides carinatus (TERQUEM); Y. LE C ALVEZ, Revision des Foraminiferes .. . , p. 45 , PI. 4, figs. 72-74.1961. Cibicides carinatus (T ERQUEM) ; J . P . H . K AASSCHIETER, Foraminifer a of the Eocene.. ., p. 221, PI. 14, fig. 6a-c.

Material. - About 30 specimens, in most cases badly preserved .Dimensions of an average specimen (in mm):

F. IX jlO40·350·320·12

Description. - Test piano-convex, completel y flat on the dorsal side, slightly convexventrally , generally somewhat lobulate and irregularly ovate in the outline, periphery stronglyacute , with a wide, angled , serrate keel. Chambers gently inflated on the ventral side, on thedor sal side arranged into at least two whorl s, the last one cont aining about 6 chambers, con­siderably increasing in size as added. Sometimes both sides are evolutely coiled. Sutures limbate,flush with surface .or indistinct, rather depre ssed on dorsal side, distinctly depre ssed on ventralside. Surface smooth on ventral side, slightly rough and coarsely perforated on dorsal side.Aperture, slit-like on the ventral side at the base of the last formed chamber.

Fi g. 16Individu al variat ion of Cibicides carinatus (TERQUEM); Pamietowo boring, 257 m, Montian.

Variation significant; it applies both to the size and to general shape of test. Specimenscan be twisted to a different degree, with a more distinct or less distinct arrangement of chambersand with differently developed peripheral margin, which is more keeled or less, serrate or smooth(see Text-fig. 16).

Remarks. - Specimens described as Cibicides carinatus from Poland do not seem todiffer from those described by TERQUEM (1882,) revised by LE CALVEZ (1949), and from tho seby KAASSCHIETER (1961) or TEN DAM (1944), all from the Eocene of Europe. In comparisonwith the specimens from France and Belgium, our s are never so pitted on the ventral side andthey are generally less lobulate in outline .

Occurrence. - In Poland common from the uppermost Maastrichtian up to the Paleocene,i. e. Montian. Present in the Paleocene , i.e. Thanetian of France (Paris Basin) and of England.Known to be common in the Eocene of France, Belgium and Holland.

76 KR YSTYNA POZAR YSK A & J ANI N A SZCZECHURA

Cibicides cuvillieri ROUVILLOTS, 1960

(PI. XII , figs. 1-7)

1960. Cibicides cuvillieri ROUVI LLOIS ; A . ROUVI LLOIS, Le T hanetien. .. , p. 74, PI. 3, fig. 51 a-c (non PI. 3, fig. 52; PI. 4,fig. 54 a-c).

1960. Cibicides subinvolutus ROUVILLOIS; A. ROUVI LLOIS, lbid., p . 73, PI. 4, fig. 53a-c.1965. Cibicides er. cryptontphalus hercegovinensis DE WITT PUYT; K. POZARYSKA, F oramini fer a and bio st ratigraphy.. ..

p. 130, PI. 26, fig. I a-c.

?1965. Cibicides sp . 2; B. MCG owRAN, Two Paleoeene foraminifcral faunas.. ., p. 55, PI. 4 , fig. 2.

Material. - Over two hundred specime ns, mostly well pre served.Dimensions of 3 specimens (in mm):

Longest d iameter

Sh ortest diamete rHeight of test

F. IX/2240·200·15

F. IX/2270·370·27

0·15

F. IX /2300-400·27

Description. - Te st piano-convex, generally co ncave on the dorsal side, convex on theventral. Outline rounded or gently oval , sometimes lobulate, peripheral margin acute, bluntlykeeled; so metimes th e keel is visible onl y on the dorsal side, not seen on the ventral one. Dorsalside somewhat evolute, early whorls forming a small central plug finely dropped , because thefinal whorl strongly embraces the preceding ones. Chambers di stinct, 8-11 in number in thelast whorl , flat on the dorsal side. di stinctly inflated on the ventral one. Sutures moderatelycurved on the dorsal side, limbate, visible between all the chambers, sometimes in the earlyportion of test onl y, and th en becoming o bscured in th e last portion; sutur es on the ventralside more radial , commonly thickened in early portion of test , but depressed between two orthree last chambers. Surface smooth, perforated on both sides, except for the raised sutures.Aperture, slit-like at the ba se of the last formed chamber, extending on the dorsal side at theinner margin along the last few chambers.

Variation considerable; it applies mainly to the size, shape and ornamentation. Thereare specimens with the outline lobulate, others not lobulate but then keeled. Chambers inflatedto different degrees, sutures more depressed or less, even distinctly limbate, especially on thedorsal side where they form roll-like thickenings around the inner whorl. There are, however,intermediate specimens, the systematic po sition of which cannot be determined as certain .

Remarks. - Specimens from Poland included into Cibicides cuvil/ieri ROUVILLOTS (1960) ,because of their great variability, probably represent a group of species clo sely related to eachother. Some of them do not differ from specimens described by ROUVILLOIS as C. cuvillieri,o r from those assigned by the same author in 1960 to C. subinvolutus, both described from theThanet bed s of the Paris Basin. Specimens assigned by ROUVILLOTS as C. subinvolutus seem,however. to rep resent less developed, probably younger specimens of C. cuvillieri species.Some of the Poli sh specimens are, on th e other hand, simila r to certain specimens of C. 10­batulus WALKER & JACOB (1798) emend. D'ORBlGNY (1846). The variability of C. lobatuluswas given earlier in BATJES paper (1958, p. 153). The Poli sh specimens , in comparison with thosereferred to C. lobatulus, are usually more clo sely a nd regularly coiled. It is possible that Ci­bicides sp. 2. described by M CGOWRAN from Australian Paleocene, represents also the abovedi scussed species, differing only slightly on the dorsal side. General appearance of specimensreferred to C. cuvillieri is also similar to that of specimens described as C. cantii by HAYNES(1957, p. 50, PI. 5, figs. 13, 13b; PI. 6, fig. 2). from the Thanetian of England. The specimens,

rORAMJNIFERA FROM TH E PALEOC EN E Or POLA ND 77

assigned by POZARYSKA (1965) as Cibicides cf. hercegovinensis cryp tomphalus DE WITT PUYT,are reclassificated in the present paper as C. cuvillieri ROUVILLOIS.

Occurrence. - In Poland, very common in the Paleocene, i. e. Danian and Montianeverywhere in the Polish Lowlands and Pamietowo boring. Known from the Paleocene, i. e.Montian of Holland (Bunde) and Belgium ("Puits Artesien" in Mons) and from the Thanetianof France (Paris Basin); probably present al so in the Paleocene of Australia.

Cibicides lectus VASILENKO, 1950

(P I. X , figs. 3-7)

1950. Cibicides (Cibicidoides) leetus VA SILENKO ; V. P. V ASILENKO, Foraminifery paleocena. .. , p. 218, PI. 6, figs. 2-3.1965. Cibicides leetus V ASILENKO ; K . P OZ ARYSKA, Foraminifera and biostra tigraphy. .. . p. 132. PI. 25, fig. 2a-e (here

additional synonymy included) .

Material. - Some hundred specimens well pre served .Dimensions, description and remarks are given in POZARYSKA'S paper (1965).

Variation considerable; it applies mainly to the size and to the development of centralpart of dorsal side. The outline is rounded or lobulate, peripheral margin transparent or not.In small, probably young specimens, the thickenings and additional apertures on the dorsalside are not yet as well developed as in large, probably adult form s (see PI. X, figs. 3-7).

Occurrence. - In Poland very common in the Paleocene, i. e. Montian of the PolishLowlands and Pamietowo boring. Known to be common in the Paleocene, i. e. Selandianof Denmark, Sweden (writers' own ob servation), U .S.S. R. (Russian Platform) and probablyin France.

Cibicides proprius (BROTZEN, 1948)

(PI. XVI , figs. 1-4)

1948. Cibicides proprius BROTZEN ; F . BROTZ EN, The Swedish Pa1eocene..., p. 78, PI. 12, figs. 3-4.1960. Cibicides ( Cibicidoides) proprius (BROTZEN) ; A. ROUVILLOIS, Le Thanet ien.. ., p. 74, PI. 4, fig. 55a-e.

?l960. Cibicides breslesensis ROUVILLOIS ; A. R OUVILLOIS , Ibid., p. 75, PI. 4, fig. 56a-e.1961. Cibicides proprius (BROTZ EN) ; J. P . H . K AASSCHJETER, Foraminifera of the Eocene , p. 229, PI. 13, fig. 9a-c.1965. Cibicides proprius (BROTZEN) ; K. P OZARYSK A, Foramin ifera and biostrat igraphy , p. 134, PI. 27, figs. I a-e,

5a-c (here additional synonymy included) .

Material. - Over one hundred specimens well preserved.De scription, variation and remarks are given in POZARYSKA'S paper (1965). It must be

added, however, that Cibicides breslesensis ROUVILLOIS is tentatively referred here to C. proprius.In the true Montian of Poland there are specimens of the same size and general appearanceas C. breslesensis, having however not so thickened sutures on the ventral side. A comparisonof C. proprius (BROTZEN) and C. breslesensis ROUVILLOIS is based on comparative material fromthe Thanetian of France, offered kindly by Mi ss A. ROUVILLOIS, and those from Selandian ofSweden sent kindly by Dr. F. BROTZEN. The specimens from France and these from Polandare both markedly smaller than specimens of typical C. proprius BROTZEN from Sweden (typelocality), but as may be observed in the Pamietowo boring, the specimens of C. proprius havechanged their size in geological time , being large in Selandian, but small in true Montian.Similarly small specimens of C. proprius occur in the Montian of Belgium ("Puits Artesien"

78 KRYSTYNA PO:lAR YSKA & JANI NA SZCZECHURA

in Mons) and Holl and (Bunde's boring) according to the writers' own ob servat ions. Thedifference between C. breslesensis ROUVILLOIS and C. proprius BROTZEN given by ROUVlLLOIS(1961) fall within the variation range of C. proprius (see PI. XVI, figs. 1--4).

Occurrence. - In Poland, common in the Paleocene, i. e. Danian and Montian of thePoli sh Lowlands and Pamietowo boring. Present in the Paleocene, i. e. Danian and Selandianof Sweden, Denmark , north and sou th regions of U.S. S. R. (Russian Platform). Known alsofrom the Paleocene, i. e. Montian of Holland, Montian and Thanetian of France (Paris Basin)and Thanetian of England.

Cibicides sahlstroemi BROTZEN, 1948

(Text-fig. 17)

1948. Cibicides sahlstriimi BROTZEN; F. BROTZEN, Th e Swed ish Paleocene . . ., p . 85, PI. 17, fig. I.1961. Gavelinella sahlstriimi (BROTZEN) ; J. HOFKER, Fo ra minifera from the Cretaceous , LIII , p . 66, fig. 5a-c.1965. Cibicides sahlstroemi BROTZEN; K. POZARYSKA, Fo ra minifera and biostratigraphy , p, 134, PI. 27, fig. 2a-c.1966. Cibicides sahlstroemi BROTZEN; K . POZARYSK A, Th e Cretaceous-Tertia ry bo undary , p. 63.

Fi g. 17Individual variation of Cibicides sahlstroemi BROTZEN; Pamietowo boring. 257 m. Montian .

Material. - One hundred specimens, mostly well preserved.Dimensions, description and remarks are given in POZARYSKA'S paper (1965).

Variation insignificant; it applies mostl y to the dorsal side ornamentation, which canbe more regular or less ob scuring the chamber arrangement (see Text-fig. 17). A very dis­tinctive species.

Occurrence. - In Poland very common from the Upper Maastrichtian up to the Paleocene(Montian) of Poli sh Lowlands and Pamietowo boring . Common in the Paleocene, i.e. Danianand Selandian of Sweden and Denmark, Montian of Belgium ("Puits Artesien" in Mons)and Holland (Bunde), and in the Paleocene, i. e. Thanetian of England.

Cibicides simplex BROTZEN, 1948

1948. Cibicides simplex BROTZEN; F . BROTZEN. Th e Swed ish Paleocene.. .. p. 83. PI. 13. fig. 4.1961. Gavelinella simplex (BROTZEN); J. HOFKER. Foraminifera from the Cretaceous LIII. p . 66, fig. 4a-c.1965. Cibicides simplex· BROTZEN; K . POZARYSKA. Foraminifera and bio stratigraphy , p. 135. PI. 26. fig. 2a-c (here

additional synonymy included) .1966. Cibicides simplex BROTZEN; K . POZARYSKA, The Cr etaceous-Tertiary boundary. ... p. 63.

Material. - Some hundred specimens well preserved .Dimensions, description and remarks are given in POZARYSKA'S paper (1965).

FORAMINIFERA F ROM TH E PALEOCEN E OF POLAND 79

Variation extremely insignificant. Thi s is one of the most uniform species .

Occurrence. - In Poland common in the uppermost Cretaceous and Paleocene, i. e. Da­nian and Montian of Polish Lowlands and Pamietowo boring. Common in the Paleocene,i.e. Danian and Selandian of Denmark and Sweden, in the Paleocene (Montian) of Holland,and in the Paleocene, i. e. Thanetian of France (Paris Basin).

Cibicides succedens BROTZEN, 1948

(Tex t-fig. 18)

1944. Cibicides cryptomphalus (REUSS) var. hercegovinensis DE W IIT PUYT ; A. TEN DAM, Die stra tigraphische... , p. 132,PI. 5, fig. 9 (non Cibicides cryptomphalus (REUSS) var. hercegovinensis DE WITT PUYT, 1941).

1948. Cibicides succedens BROTZEN; F. BROTZEN, The Swedi sh Paleocene. .. , p. 80, PI. 12, fig. 2.1965. Cibicides succedens BROTZEN; K. POZARYSKA , Foraminifera and biostrat igraphy. . ., p . 136, PI. 28, figs. I , 5 (here

additional synonymy included).

® ..,,'"''''''''". "..J".".--:~

"'~ "" ......' ~-" --Ib

Fig. 18Individual vari ation of Cibicides succedens BROTZEN ; Parn ietowo boring , 231 m, Montian.

Material. - Some hundred specimens well preserved.Dimensions and description are given in POZARYSKA'S paper (1965).

Variation concerns the general shape of the test and development of central plug onventral side, which is figured above (see Text-fig . 18).

Longest diameterShortest diam eterHeight of test

~o KR YSTYNA POZ AR YSK A & JA NI NA SZCZECH URA

Remarks and comparison with other species are given in detail by BROTZEN (1948).

Occurrence. - In Poland common in the Paleocene (Montian) of Poli sh Lowlandsand Parnietowo boring. Known as common in all the Paleocene of Europe.

Genus PLANULINA [)'ORBIGNY, 1826

Planulina Iimbata BROTZEN, 1948

(PI. XI, fig. 2)

1948. Planulina limbata BROTZEN ; F. BROTZEN, Twe Swedish Paleocen e. .. , p, 85, PI. 13, fig. Ta-c.1962. Discopulvinulina trinitatensis CUSIIMAN & RENZ; J . HOFKER , Foraminifera from the Cretaceous.. ., LVII , p. 8,

fig. 1a-c (non Discorbis midwayensis CUSHMANvar . trinita tensis CUSHMAN & RENZ; J . A. CUSHMAN (1951), PaleoceneForaminifera. .. , p. 48, PI. 13, fig. 22a- c).

non 1963. Gavelinella limbata (BROTZEN); J . HOFK ER, Foraminifera from the Cret aceous.. ., LXIII, p. 6, fig. 2a -c.

Material. - A few specimens, generally damaged .Dimensions of one average specimen (in mm) :

F. IXjll 90·300·250·12

Description. '- Te st small, trochospiral , stro ngly compressed , only slightly convex onthe dorsal side, concave on ventral side, ovate and gently lobulate in outline. Periphery ratheracute, with a limbate border as a continuation of th e rai sed spiral and septal sutures of the dorsalside. Chambers coiled in 2-3 whorls visible on dorsal side, only the last one visible ventrally.In the last whorl, there are about 6 chambers moderately increasing in size as added. Umbilicusshallow, filled by a plug or open. Sutures limbate and curved on the dorsal side, somewhatdepressed on the ventral. Surface finely porous. Aperture slit-like on the ventral side, runningfrom umbilicus to margin under a small lip , extending as far as 3 last chambers.

Variation not studied due to the scarcity of material.

Remarks. - Our specimens referred to Planulina limbata differ from specimens describedunder this name by BROTZEN (1948) only in having less inflated chambers. They do not differ,however, from the specimens described by HOFK ER (1962) from the lowermost Tertiary ofHolland, determined as Discopulvinulin a trinitatensis (CUSHMAN & R ENZ). Specimens from thefillings of the holes in the hard ground in Curfs quarry (Holland) and the Paleocene (Selandian)of Denmark, included into Gavelinella limbata by HOFKER (1963) , differ greatly from thosedescribed by BROTZEN from the Paleocene of Sweden, and probably represent other species .In contrast with holotype, HOFKER'S specimens have a broadly rounded peripheral marginand their chambers are much more inflated. Our specimens are simila r in general appearanceto those described by VAN BELLEN (1946) as Discorbis plana from the Eocene (recte Montian)of Holland. VAN BELLEN'S illustrations are not sufficient ly clear, but it seem s that the figuredspecimens have less limbate and less curved sutures not rai sed at all , than those of BROTZEN'Sspecies from Sweden and Poland.

Occurrence. - In Poland very rare in the Paleocene (Montian) of Pamietowo boringonly. More common in the Paleocene, i. e. Selandian of Sweden, Denmark and in Montianof Belgium ("Puits Artesien" in Mons) and Holland (Bunde).

FORAMINIFERA FROM TH E PALEOCENE OF POLAND

Family NONIONIDAE SCHULTZE, 1854

Genus NONION DE MONTFORT, 1808

Nonion graniferum (TERQUEM. 1882)

(PI. IX, figs. 10-12: Text-fig. 19)

81

1882. Nonionina granifera TERQUEM ; O. TERQUEM, Les Foraminiferes de l' Eocene . .. , p, 42, figs. 8a-b , 9a-b .1965. Nonion graniferum T ERQUEM ; K . POZARYSKA . Foraminifera and bio str atigraphy.. ., p, 93. PI. 21, fig. 5a-b (here

additional synonymy included) ,

Fig. 19Individual variation of Nonion graniferum (TERQUEM) ; Pamietowo boring, 257 m, Montian.

Material. - About one hundred specimens well preserved.Dimensions, description and remarks are given in POZARYSKA'S paper (1965).

Variation not significant; there are specimens of different size and number of chambers.Their umbilical region as well as sutures and ornamentation are inequally developed, as isseen on figures above (see Text-fig. 19).

Occurrence. - In Poland common in the Paleocene, i. e. Danian and Montian of PolishLowlands and Pamietowo boring. Known from the Paleocene (Selandian) of Sweden andDenmark, Paleocene of Asia and America, Paleocene (Thanetian) and Eocene of France(Paris Basin).

Nonion sublaeve TEN DAM, 1944

(PI. IX, figs. 13-15)

1944. Nonion sublaeve TEN DAM ; A. TEN DAM, Die stratigraphische. .. , p, 109, PI. 3, fig. 8a-b.1956. Protelphidium sublaeve (TEN DAM) ; J. HAYNES, Certain smaller British ..., p. 87, PI. 16, figs. 2, 2e.1960. Protelphidium sublaeve (TEN DAM); A. ROUVILLOIS, Le Thanetien. . ., p, 65, PI. 2, fig. 23.

Pal ueont o lo gia Polon ica No 20. 6

82 KR YSTYNA POZAR YSKA & JANINA SZCZECH URA

Material. - About 30 specimens well preserved.Dimensions of 2 specimens (in mm):

Longest diameterShortest diameterHeight of test

(F. IX/58)0·320·28

(F. IX/59)0·330·280·09

Description. - Test planispiral and involute , equally compressed on both sides, slightlylobulate in outline, periphery broadly rounded. Chambers 9 in number, increasing graduallyin size as added. Sutures depressed, slightly curved, enlarging at the umbonal region; umbonalregion rather large, covered by a distinct , transparent, more regular or less regular knob andby a more numerous or less numerous fine protuberances. Surface smooth, glossy. Aperturegenerally obscured, but sometimes a row of small apertural pores is seen at the base of the lastformed chamber.

Variation insignificant , concerning mainly the size and the general appearance of test,especially in the outline. Somet imes, a central knob in the umbonal area is not developed orreduced.

Remarks. - Specimens referred to Nonion sublaeve from Poland do not differ fromthose, described as N. sublaeve by TEN D AM (1944) from the Paleocene of Holland.

Occurrence. - In Poland, not common in the Paleocene, i.e. Montian of Pamietowoboring only. Known from the Paleocene of Holland, and from the Paleocene , i.e. Thanetianof France and England.

Genus NONIONELLA CUSHMAN, 1926

Nonionella communis communis (D'ORBIGNY, 1846)

(PI. IX. fig. 6)

1846. Nonion communis O'ORBIGNY ; A. O'ORBIGNY. Forarniniferes fossiles.. .. p. 106. PI. 5. fig. 7 ; fide Cat. of Foram.,ELLIS & MESSINA.

1882. Nonion commune (O'ORBIGNY) ; O. TERQUEM, Les Fo ra miniferes de l'Eocene.... p. 42. PI. 2. fig. 6.1939. Nonion commune (O'ORBIGNY); J. A. CUSHMAN, A Monograph of the foraminiferal famil y.... p. 10, PI. 3, fig. 2.1944. Nonion acutidorsatum TEN D AM; A. TEN DAM. D ie stratigra phische..., p. 108, PI. 3. fig. 19a-b.1950. Nonion commune (O'ORBIGNY); Y. LE CALVEZ, Revision des Foraminiferes.... p. 52.

Material. - A dozen or so specimens well preserved.Dimensions of one specimen (in mm):

Longest diameterShortest diameterHeight of test

F. IX/2230'460'320·25

Description. - Test indistinctly trochospiral, nearly planispiral , strongly compressed,coiled asymmetrically and having the last chamber slightly overhanging the former ones. Outlineovate , indi stinctly lobulate, peripheral margin rather angled . Chambers low, triangular, 9 innumber in the last whorl , rapidly increasing in size as added, sutures gently curved, very slightlydepressed, nearly flush with surface. Surface smooth, glossy. Aperture interiomarginal nearthe periphery, extending on the ventral side.

FORAMINIFERA FROM THE PALEOCENE OF POLAND 83

Variation insignificant; it concerns mainly the size and the degree of compression of test.

Remarks. - Our specimens are very simila r to the figured holotype of Nonion communis(recte Nonionella communis communis ) D'ORBlGNY, differing only in having the peripheralmargin not so angular. The differences with a new erected subspecies Nonionella communispaleocenica are given below.

Occurrence. - In Poland. not common in th e Paleocene (Montian) of Parnietowo boring.Known from the Paleocene (Montian) of Holland and Eocene of France (Pari s Basin) .

Nonionella communis paleocenica n. subsp.

(PI. IX. figs. 7-9)

Holotypus: Specimen present ed 0 11 PI. IX, fig. 9 (F. IX/226) .Stra tum typicum: Monti an.Locus typicus: Pami ctowo boring, depth 208·5 m (North Poland).Derivalio nominis: Lat . paleocenica = common in the Paleocene beds.

Longest diameterShortest diameterHe ight of test

Material. - About 40 specimens well preserved .Dimensions of two specimens (in mm):

F. lX/2260·440·270·20

F. IX j2240-400·25

Description. - Te st subtro choida l, compressed , ovate in outline. Dorsal side partiallyevolute with an umbonal bo ss, ventra l side completely involute. Peripheral margin slightlyrounded. Chambers strongly increasing in size a s added, inequilateral, especially the last one,about 9 in number in the la st whorl. Sutures radial or slightly curved, depressed between thelast chambers, flush with surface in the earlier portion of te st. Surface smooth, except for theumbilical depression and umbonal boss, which are covered by small tubercles, sometimeswedging in between the inner parts of chambers. Aperture interiomarginal, slit-like near theperiphery, extending slightly onto the ventral side.

Variation not significant, concerns the size of test, as well as the coiling of whorls. Thereare specimens with more distinct or less di stinct umbonal bo ss, which is not always coveredby ornamental tubercles.

Remarks. - Our specimens are the most similar to those referred to Nonionella communiscommunis (D'ORBIGNY, 1846), they differ, however, in being more trochospiral, not so closelyinvolute and in having a distinct umbilical depression, generally filled up with more numerousor less numerous small tubercles.

Occurrence. - In Poland. common in the Paleocene (Montian) of the Pamietowo boringonly.

Nonionella ovata BROTZEN, 1948

pars 1926. Nonionina turgida PLUMMER (no t WILLlA MSON) ; H . J . PLUMMER, Foraminifera of the Midway.... p. 159,PI. 12, fig. 7.

1948. Nonionella ovata BROTZEN; F. BROTZEN, The Swedish Paleocene.. ., p. 68, PI. 10, figs. 13-14.1951. Nonionella sp.; J. A. CUSHMAN, Paleocene Foraminifera.. .. p. 37, PI. 11, figs. 5-6.1965. Nonionella ovata BROTZEN; K. POZARYSKA. Foraminifera and biostratigraphy.. ., p , 94, PI. 21, fig. 1a-c.

6*

84 KR YSTYNA POZARYSKA & JAN1NA SZCZECH URA

Material, - Over one hundred specimens, mo stly well preserved.Dimensions, description, va ria tion and remarks are given in BROTZEN'S (1948) and in

POZARYSKA'S (1965) papers.

Occurrence, - In Poland, common in the Paleocene (Montian) of Pamietowo boringand the Poli sh Lowlands. Known to be seldom in Paleocene (Selandian) of Sweden and Denmark.

Family ALABAMINIDAE HOFKER, 1951

Genus ALABAMINA TOULMIN, 1941

AIabamina midwayensis BROTZEN, 1948

(Text-fig. 20)

1948. Alabamina midwayensis BROTZEN; F. BROTZEN, The Swedish Paleocene.. ., p. 99, PI. 16, figs. 1-2.1965. Alabamina midwayensis BROTZEN; K . POZARYSKA. Foraminifera and biostratigraphy. .. , p. 114, PI. 21, fig. 3a-c

(here additional synonymy included) ,

Material, - Som e hundred specimens well preserved.Dimensions and description are given in POZARYSKA'S paper (1965). Variation discussed

by BROTZEN (1948) and HOFKER (1960).

Remarks. - Within specimens found in the Montian of Pamietowo boring, there aresome with a more convex dorsal side, than the true Alabamina midwayensis, and they seem torepresent rather A. wilcoxensis described by TOULMIN (1941) from the Wilcox formation ofAlabama, U. S. A. (see Text-fig. 20).

F ig. 20Ind ividu al variation of Alabamina midwayensis BROTZEN; Pamietowo boring, 231 m, Montian.

Occurrence. - In Poland, very common in the uppermost Maastrichtian and Paleocene,i.e. Danian and Montian of Poli sh Lowlands and Parni etowo boring. Known to be commonin the Paleocene, i. e. Danian and Selandian of Sweden, Denmark, Montian of Holland andPaleocene of Egypt. Pr esent in the Paleocenc, i. e. Th an etian of France (Paris Basin) and England.Present also in U. S. S. R . (Russian Platform and Asiatic part).

FORAMINI FERA FROM TH E PALEOCENE OF POLAND

Family OSANGULARIIDAE LOEBLlCH & TAPPAN, 1964

Genus GYROIDINOIDES BROTZEN, 1942

Gyroidinoidcs octocamerata (CUSHMAN & HANNA, 1927)

85

1927. Gyroidina soldanii U'ORBIGNY subspecies octocamerata CUSHMAN & HANNA; J . A. CUSHMAN & C. D. HANNA,Foram inifera from the Eocene. . ., p, 223, PI. 14, figs. 16-18.

1965. Gyroidinoitles octocamerata CUSHMAN & HANNA; K. POZAR YSKA, Foraminifera and biostrat igraph y. . ., p. 107,PI. 18, figs. 3a-c, Sa-c (here additional synonymy included) .

1965. Gyroidinoides octocamerata CUSHMAN & HANNA; B. MCGOWRAN, Two Paleocene . . ., p. 47, PI. 2, fig. 5.

Material. - Over two hundred specimens well preserved.Dimension s, description , variation and remarks are given in POZARYSKA'S paper (1965).

Occurrence. - In Pol and very common from the upperm ost Maastrichtian up to theMontian of Polish Lowlands and Parnietowo boring. Known as common in the Upper Ma­astrichtian a nd Paleocene, i. e. Danian and Montian of Europe and Australia, and the Eoceneof America.

Gyroidinoides pontoni BROTZEN, 1948

1948. Gyroidinoides pontoni BROTZEN; F. BROTZEN, The Swedish Paleoccne... , p. 76, PI. 11, figs. 4, 5.1965. Gyroidinoides pontoni BROTZEN; K. POZARYSKA, Fora minifera and biostratigraphy. .. , p. 108, PI. 18, fig. 4a -c

(here additional synonymy included) .

Material. - Over two hundred specimens well pre served.Dimensions, description, variation and remark s are given in POZARYSKA'S paper (1965).

Occurrence. - In Poland very common in the Paleocene, i. e. Danian and Montian(Selandian and true Montian) of Poli sh Lowl ands and Pamietowo boring. Known as commonin the Paleocene of Sweden , Denmark and Belgium .

Genus OSANGULARIA BROTZEN, 1940

Osangularia cordieriana navarroana (CUSHMAN, 1938)

1938. Pulvinulinella navarroana CUSHMAN; J. A. CUSHMAN, Some new species.. ., p, 66, PI. It , fig. 5.1962. Osangularia cordieriana navarroana (CUSHMAN, 1938) ; E. V. HERMANNI, Zu r Artfa ssung von Osangularian .. ..

p. 280, PI. 19, figs. 2-4.1965. Osangularia cordieriana navarroana (CUSHMAN); K . POZARYSKA, Foramini fera and biostratigraphy.. ., p. 112,

PI. 19, figs. I a-c. 5a-b (here additional synonymy included),

Material. - Some hundred specimens well pre served.Dimensions, description, variation and remarks are given In POZARYSKA'S paper (1965) .

Occurrence. - In Poland very common in the Paleocene, i.e. Danian and Montian ofPolish Lowl ands; in the Pamietowo boring only a few specimens just a bove the Maastrichtian.Common in the Paleocene of Sweden, Denmark, U. S. S. R. (Russian Platform) and Holland,where it starts from the uppermost Maastrichtian . It is not ob served in the true Montian strata.Probably pre sent in the Paleocene, i. e. Thanetian of France (Pari s Basin) and England.

86 KRYSTYNA POZARYSKA & JANINA SZCZECHURA

Family ANOMALINIDAE CUSHMAN, 1927

Genus ANOMAUNA O'ORBIGNY, 1826

Anomalina burlingtonensis (JENNINGS, 1936)

(PI. XVI , figs. 5-6)

1936. Cibicides burlingt onensis JENNINGS; P. H . JENNI NGS. A microfauna.. ., p. 39, PI. 5, fig. Sa-c.1965. Anomalina burlingtonensi s (JENNINGS), K. POZ AR YSK A, Foraminifcra and biostratigraphy. . ., p. 127, PI. 26, figs. 3-4

(here additional synonymy included) .

1966. Cibicides burlingtonensis JENNINGS; J. HOFKER, Maestricht ian , Oanian and Paleo cenc ... , p. 239, PI. 47, fig. 21 ;PI. 48, fig. 54.

Material. - About one hundred specimens, in most ca ses well preserved.Dimensions, description, variation and remarks are given in POZARYSKA'S paper (1965);

here ought to be added , however, that specimens from Poland , referred to this species, seemto be very similar to compressed specimens of Anomalina danica (BROTZEN); more than onceit is difficult to distinguish with certainty small , compressed representatives of A. danica andthese referred to A. burlingtonensis.

Occurrence. - In Poland common in the Paleocene, i. e. Danian and Montian, in thePolish Lowlands, as well as in Parnietowo boring. Known from the uppermost Cretaceous upto Paleocene in Limburg, Paleocene, i. e. Danian and Selandian of Sweden and Denmark,Paleocene of U. S. S. R. (Asiatic part) and Eocene of North Am erica.

Anomalina danica (BROTZEN, 1940)

(PI. XIV , figs. 6-11)

1940. Cibicides danica BROTZEN; F. BROTZEN, Flintriinna ns... , p. 31, PI. 7, fig. 2.1948. Anomalino ides danica (BROTZEN); F. BROTZEN, The Swedish Paleocene ... , p, 87, PI. 14, fig. I; Text-fig. 22.1965. Anomalina danica (BROTZEN) ; K. POZARYSKA, Foram inifcra and biostrat igraph y.. ., p. 128, figs. 1-3 (here additional

synonymy included).1966. Gavelinella danica (BROTZEN) ; J. HOFKER, Maestri chtian, Danian and Paleocene .. ., p. 227, PI. 43, fig. 86 a-c.

Material. - Some hundred specimens very well preserved.Dimensions of 3 specimens (in mm):

Longest diamet erShortest diameterHeight of test

F. IX/2270·300·25

F. IX/2300·520-45

F. IX/23 I0·900·650·19

Description and variation are given in BROTZEN'S (1940, 1948), HOFKER'S (1955, 1966)and POZARYSKA'S (1965) papers.

Remarks. - Our specimens are described as Anomalina danica BROTZEN, because inmost cases they do not differ from specimens of thi s species described from Sweden. They alsoseem to be very similar to forms occurring in higher strata in Europe, assigned either toA. granosa (VAN BELLEN, not HANTKEN) or to A. grosserugosa GUMBEL. Undoubtedly, theyall are related to each other, being changed in geological time only very slightly, especiallyin ornamentation. The variation within our rich collection confirms thi s opinion. Variationis significant, applying to several features such as size, general shape and ornamentation.

FORAMINIFERA FROM THE PALEOCENE OF POLAND 87

Specimens are peripherally lobulate to different degrees , having chambers more inflated or less,with sutures distinctly depre ssed, flush with surface or strongly thickened. The umbilical de­pression is filled or open , forming a distinct hollow extending along the last formed chambers.Surface of test is nearly smooth, generally fairly pitted or sharply rough , even granulated. Thereare specimens distinctl y compressed, finely perforated, resembl ing specimens referred toA . burlingtonensis JENNINGS; both these species coexist in many samples of Polish Montian.

Occurrence. - In Poland very common in the uppermost Maastrichtian up to the Pa­leocene, as in the Lower Tertiary sediments all over the world .

Anomalina ekblomi (BROTZEN, 1948)

(PI. XlII , figs. 1-4)

1948. Cibicides ekb lomi BROTZEN; F. BROTZEN, The Swedish Paleocene..., p. 82, PI. 13, fig. 2a- c.?1965. Cibicidina mariae (RUPERT JONES); B. McGowRAN, Two Paleocene..., p. 51, PI. 4, fig. I.1965. Anomalina ekb lomi (BROTZEN); K. POZARYSKA, Foraminifera and biostratigraphy..., p, 127, PI. 27, fig. 6 (here

additional synonymy included).1966. Gavelinella ekblomi (BROTZEN) ; J. HOFK ER, Maestrichtian, Danian and Paleocene..., p. 197, PI. 36, fig. 26 ; p. 208,

PI. 39, fig. 44; p . 259, PI. 56, fig. 108.

Ma terial. - Over three hundred specimens well preserved.Dimensions, description and variation are given in POZARYSKA'S paper (1965).

Remarks. - In the MCGOWRAN'S opinion (1965) , Cibicidina mariae (RUPERT JONES)is conspecific with Anomalina ek blomi (BROTZEN). Taking under consideration both illustrationsof specimen of C. mariae from Paleocene of Australia given by Mc GOWRAN, and specimenskindly sent us by HAYNES from Thanetian of England (from where is described C. mariae) ,it results that C. mariae have less chambers and generally is more inflated than A. ekblomispecimens.

Occurrence. - In Poland very common from uppermost Maastrichtian up to Montianof Polish Lowlands and in Parnietowo boring. Known to be common in the Paleocene (Se­landian) of Denmark, Sweden, Austr ia, Germany, U. S.S. R. (Russian Platform) and Australia.Present in the Paleocene, i.e. Thane tian of England and France (Paris Basin).

Anomalina minor D. sp.

(PI. XIV, figs. t -5 ; Text-fig. 21)

Holotypu s: Specimen presented on PI. XIV, fig. 3 (F . IX/235).Paratypi: Specimens presented on PI. XIV, figs. I, 2, 4 (F. IX/233, 234, 236).Strati/m typicum: Montian.Locus typicus: Pamietowo boring, depth 205 m (North Poland).Derivatio nominis: Lat. minor = smaller; because of its small size in comparison with other Anoma/ina species.

Ma terial. - About 30 specimens well preserved.Dimensions of 4 specimens (in mm):

Longest diameterShortest diameterHeight of test

F. IXj2330·200·15

F. IX/2340·250'22

F. IX/2350·300·220·16

F. IX/23 60·160'12

88 KR YSTYNA POZAR YSKA & JANINA SZCZECHURA

Description. - Test small, almost planispiral , rather compressed, gently lobulate andovate in outline, periphery broadly rounded. Chambers slightly inflated, arranged in at leasttwo whorls, the outer one containing up to 5-6 chambers, moderately increasing in size asadded. Sutures curved , somewhat depressed on both sides, those between the earlier portionof the test are completely ob scured on the ventra l side. The inner coil on ventral side is partlybordered by a deep furrow, extending on the apertura l area along the ba se of the last chamber.Aperture slit-like, marginal, extending somewhat onto the ventral side. Surface strongly pitted,but only on 2-3 last chambers on ventra l side, slightly porous on dorsal side ; sur face of olderchambers, especially on ventral side con spicu ou sly smooth, glistening.

~: .... '.: ...

•. "0.·

" .... .....' ..

<I .

;)

(!)..... ..... .. .... .

' .

s

Fig. 21Anomalina minor n, sp. ; / --4 microspheric for ms, 5-8 megalosphcric forms; Pamietowo bor ing, 257 m, Montian .

Variation insignificant ; it applies in the first place to the size and number of chambersin the last whorl and to the degree of development of surface perforation. Pits on ventral sideare strongly variable in their size and distribution. There occur micro- and megalospheric form s(see Text-fig. 21).

Remarks. - Anomalina minor n. sp. is a well distinguished species from the other describedspecies of the Anomalina genus. Some specimens of A. minor resemble mo st young specimensof A. danica BRoTzEN, from which they differ in being distinctly smaller, differently pitted andin having a characteristic furrow near the center of ventra l side.

Occurrence. - In Poland not common, in the Paleocene (Montian) of Poli sh Lowlandsand Pamietowo boring. According to the pre sent writers' observations, pre sent in the Paleocene(Montian) of Belgium ("Puits Artesien" in Mon s).

Anomalina praeacuta VASILENKO, 1950

pars 1937. Anomalina acuta PLUMMER ; M. F. GLAESSNER, Stud ien tiber Foraminiferen , p. 386, PI. 5, fig. 39.1950. Anomalina praeacuta V ASILENKO ; V . P. VA SILENKO, Foraminifery paleocena , p. 208, PI. 5, figs. 2-3.1965. Anomalina praeacuta VA SILENKO; K. P OZARY SKA, Foram inifera and biostratigraphy... , p. 129, PI. 28, fig. 2a-c

(here additional synonymy included) .

FORAMINIFERA FROM THE PALEOCEN E OF POLAND 89

Material. - Some hundred specimens well preserved.Dimensions, description, variation and remarks are given In VASILENKO'S (1950) and

POZARYSKA'S (1965) papers.

Occurrence. - In Poland very common from uppermost Maastrichtian up to the Montianin Polish Lowlands and in Pamietowo boring. Common in the Paleocene, i.e. Danian andMontian (Selandian) of Sweden, Denmark and U. S.S. R. (Russian Platform).

Anomalina umbilicata costata n. subsp.(PI. XIII, figs. 8-10)

Holotypus: Specimen presented on PI. XIII , fig. 10 (F. IX/243).Paratypi : Specimens presented on PI. XIII, figs. 8-9 (F. IX/24I, 242).Stratum typicum: Montian.Locus typicus: Pamietowo boring, depth 208 m (North Poland).Derivatio nominis: Lat. costata = with ribs; having rib-like chambers inflation.

Material. - A dozen or so specimens, well preserved.Dimensions of 3 specimens (in mm):

F. IX/241 F. IX/242 F. IX/243Longest diameter 0·17 0·22 0·35Shortest diameter 0·15 0·17 0·25Height of test 0·15

Description of this subspecies does not differ much from that given for Anomalina umbi­licata umbilicata (BROTZEN). The differences concern exclusively the development of chamberwalls, which are distinctly inflated along all sutures on both sides of test. Moreover, theperipheral margin is more acute in A . umbilicata costata n. subsp. than in A. umbilicata umbilicata.

Variation not significant. Our specimens differ mainly in size and in the degree of coiling.Some of them are closely involute , especially the small ones, others not so closely, showingslightly the inner margins of the earlier whorl s.

Remarks. - The specimens referred to Anomalina umbilicata costata n. subsp. occurmainly in the higher strata of Montian in the Parnietowo boring. Its appearance seems to bea result of an evolutionary trend of A. umbilicata umbilicata (BROTZEN).

Occurrence. - In Poland not so common, in the Paleocene (Montian) of Parnietowoboring only.

Anomalina umbilicata umbilicata (BROTZEN, 1948)

(PI. XIII, figs. 5-7)

F. IX/2390·350·25

Longest diameterShortest diameterHeight of test

Material. - About 120 specimens, in most casesDimensions of two specimens (in mm):

F. IX/2380·400·300·20

?1926. Truncatulina midwayensis PLUMMER; H. J. PLUMMER, Foraminifera of the Midway.. ., p. 141, PI. 9, fig. 7.?1948. Anomalina midwayensis (PLUMMER); F. BROTZEN, The Swedish Paleocene..., p. 88, PI. 14, fig. 3.

1948. Cibicides umbilicata BROTZEN; F. BROTZEN, The Swedish Paleocene..., p. 84, PI. 13, fig. 6.1960. Anomalina simplex (BROTZEN); E. K. SUCKAJA, Foraminifery verchnego paleocena , p. 250, PI. 3. figs. 1-2.1961. Gavelinella umbilicata (BROTZEN) ; J. HOFKER, Foraminifera from the Cretaceous LIB, p , 66, Text-fig. 6.1966. Gavelinella umbilicata (BROTZEN); J. HOFKER, Maestrichtian, Danian and Paleocene , p. 227, PI. 42, figs. 83-84.

well preserved .

90 KR YSTYNA POZARYSKA & JANINA SZCZECHURA

Description. - Test trochospiral , low, nearly planispiral, somewhat compressed, almostround and lobulate in outline, periphery broadly rounded. Chambers distinct, inflated, especiallythe last one , moderately increasing in size as added , arranged in at least two whorls , the lastcontaining 10 chambers clearly visible on the ventral side. Sutures curved, usually depressed,more on the ventral side, where chamber margins overlap each other forming a false thickening,with the exception of margin of last chambers. Umbilical depression small but distinct. Surface,when well preserved, porcellanous, smooth, finely porous. Aperture, peripheral at the baseof the last chamber, extending onto the umbilical hollow on the ventral side, with a narrowlip above. Aperture continuous beneath the inner, lip-like part of earlier chambers.

Variation not significant, concerning mainly the size and general appearance of specimens,which may be more lobulate or less in outline and may have a more developed or less aperturalregion. There are forms closely coiled or somewhat uncoiled.

Remarks. - Our specimens do not differ from the illustrated holotype of that speciesfrom Sweden. Specimens described from Limburg by HOFKER (1961) are more perforated andmore compressed. Specimens, described by SUCKAJA (1960) as Anomalina simplex BROTZENfrom the Paleocene of Crimea, are quite the same as Poli sh ones referred to A. umbilicata um­bilicata (BROTZEN). Specimens referred to A. umbilicata umbilicata (BROTZEN) from Polandare certainly very near to the specimens, figured and described by PLUMMER (1926) as Trun­catulina midwayensis (recte Anomalina midwayensis) from the Midway of America, and byBROTZEN (1948) from the Swedi sh Paleocene as Anomalina midwayensis. In contrast with theA. umbilicata umbilicata, they have more pronounced , distinctly limbate sutures. Taking underconsideration the topotypes of A . midwayensis from America and specimens referred here toA. umbilicata umbilicata from Poland, it seems the above mentioned differences could be un­derstand as connected with intraspecific variation within firstly described A. midwayensis(PLUMMER) species. That is why A. midwayensis (PLUMMER) is here tentatively included intothe synonymy of A. umbilicata umbilicata.

Occurrence. - In Poland very common in the Paleocene (Montian) in Pamietowo,Sochaczew and Boryszew borings. Pre sent in the Paleocene, i. e. Selandian of Denmark andSweden, Montian of Belgium ("Puits Artesien" in Mons), Paleocene, i. e. Thanetian of Englandand Paleocene of U. S. S. R. (Asiatic part).

Genus COLEITES PLUMMER, 1934

Coleites reticulosus (PLUMMER, 1926)

1926. Pulvinulina reticulosa P LUMMER ; H. J . PLUMMER, Forarninifera of the Midway... , p. 152, PI. 12, fig. 5.1965. Coleites reticulosus ( PLUMMER); K. POZARYSKA. Foramini fera and biostratigraphy.. ., p. 117, PI. 20, figs. t a-c,

2a-b (here additional synonymy included).

Material. - Twenty specimens usually damaged .Dimensions, description , variation and remarks are given in POZARYSKA'S paper (1965).

Occurrence. - Not common in the Paleocene (Montian) of Poli sh Lowlands and Pamie-towo boring. A very similar species occurs in the Paleocene (Montian) of France (MARtE, MS)and Belgium ("Puits Artesien" in Mons) and in the Paleocene of Sweden. Known as commonin the Paleocene, i. e. Danian and Montian and Eocene of America and Africa .

FORAMINIFERA FROM THE PALEOCEN E OF POLAND

Genus GAVELINELLA BROTZEN, 1942

Gavelinella lellingensis BROTZEN, 1948

(PI. X II , fig. 8)

91

1948. Gavelinella lellingensis BROTZEN; F . BROTZEN, The Swed ish Paleocene . . . , p. 75, PI. 2, figs. I, 2 ; Text-fig. 20.1966. Gavelinella lellingensis BROTZEN; J. HOFKER, Maestrichtian , Danian and Paleocene.. ., p . 227 , PI. 43, fig. 85a-c.

Material. - About 30 specimens, in mo st cases well preserved.Dimensions of an average specimen (in mm) :

Longest diameterShortest diameterHeight of test

F. IX /2440'500·460·26

Description. - Test trochospiral , flattened slightly on dorsal side , convex on ventralside, lobulate, nearly round in outline, peripheral margin broadly rounded. All whorls visibleon dorsal side , the inners ones indistinctly, onl y chambers of final whorl are seen around theumbilicus. Chambers gently inflat ed , increasing in size and height as added, especially so th ethree last ones on ventral side. Sutures curved more dorsally , depressed between the early cham­bers , generally thickened and transparent. Aperture a low marginal slit extending from theperiphery to the open umbillicus, bordered above by a narrow lip. Surface coarsely pitted.

Variation insignificant; it concerns mainly the size of test and the development of um­bilicu s.

Occurrence. - In Poland, not very common in the Paleocene (Montian) of Poli sh Low­lands and Pamietowo boring. Known to be common in the Paleocene (Selandian) of Swedenand Denmark; probably present also in the Montian of Paris Basin.

Genus KARRERIA RZEHAK, 1891

Karreria fallax RZEHAK, 1891

(Text-fig. 22)

1891. Karreria fallax RZEHAK ; A. RZEHAK , Di e Foraminiferenfauna .. . , p. 4, PI. 7, figs. 7a-c, 8 a-b.1965. Karreria falla x RZEHAK ; B. M CGOWRAN, Two Paleocene.. ., p. 56.1965. Karreriafallax RZEHAK ; K. POZARYSKA, Foraminifera and bio stratigraphy... , p. 138, PI. 19, figs. 3-4 (here additional

synonymy included) .

Material. - Some hundred specimens well preserved.Dimensions, description and remarks are given in POZARYSKA'S paper (1965).

Variation significa nt, presented in detail by BROTZEN (1948). It applies mainly to generalshape of test, which can be closely coiled or uncoiled to a different degree, up to rectilinear.Th e spira l side varies being flat or even concave irregularly, accord ing to the ba se on whichthe test was fixed. Moreover, th e ap erture may occur at the base of the last formed chamberor at its top, being under the shape of a slit or a round hollow, sometimes represented bya double aperture. Examples of morphological variation are given below (see Text-fig. 22).

92 K RYSTYNA POZA R YS KA & JAN INA SZCZECHU RA

(~.<~III

O.1mm

F ig. 22Ind ividual var iation of Karreria fa llax R ZEHAK ; Bo ryszew boring , 202 m, Mo ntian .

Occurrence. - In Poland very com mo n in the Paleocene, i. e. Danian and Montian ofPol ish Lowlands and Pami etowo boring. Known as co mmo n from th e uppermost Cretaceousup to the lowermost Tertiary in Europe and Au stral ia.

Ge nus PULSIPHONINA BROTZ EN, 1948

Pulsiphonina prima ( PLUMMER, 1926)""

1926. Siphonina prima P LUMMER ; H. J. P LUMMER, Fo ra minifera of the Midway.. ., p. 148, PI. 12, fig. 4.1960. Siph onina (Pulsiphonina) cf. prima (PLUMMER) ; A. R OUVILLOIS, Le Th anet ien , p. 70, PI. 2, fig. 36a-b.1961. Siphonina prima P LUMMER ; J. P. H. K AASSCHJ ETER, Foraminifera of the Eoce ne , p. 230, PI. 15, figs. 2, 15.

FORAMINIFERA FROM THE PALEOCENE OF POLAND 93

1962. Siphonina (Pulsiphonina ) prima (PLUMMER); M. E. SCHMlD, Die Foraminiferenfauna. . ., p. 344, PI. 4, fig. JI a-b.1965. Siphonina (Pulsiphonina) prima (PLUMMER); B. MCGowRAN. Two Paleocene... , p. 47.1965. Pulsiphonina prima (PLUMMER); K . POZARYSKA, Foraminifera and biostr atigraphy... , p. Ill, PI. 16, figs. 3a-c,

6a-e (here additional synonymy included) .

Material. - Over one hundred specimens, rather well preserved.Dimensions , description, variation and remarks are given in POZARYSKA'S paper (1965).

Occurrence. - In Poland common from the uppermost Cretaceous up to the Montianof Polish Lowlands and Pamietowo boring. Common in the Paleocene (Danian and Montian)of Belgium and Holland, Danian (?) of Austria, Paleocene of Sweden, Denmark and V. S. S. R.(Russian Platform), Paleocene (Thanetian) of England, and Paleocene of America (V. S. A.)and Australia.

Family CERATOBULIMINIDAE CUSHMAN, 1927

Genus CERATOBULIMINA TOULA S.S. , emend. FINLAY, 1939 and TROELSEN, 1954

Ceratobulimina tuberculata BROTZEN, 1948

1948. Ceratobulimina tuberculata BROTZEN; F. BROTZEN, The Swedish Paleocene . .. , p. 124. PI. 19, figs. 2-3.1965. Ceratobulimina tuberculata BROTZEN; K. POZARYSK A. Foraminifera and biostratigraphy. .. . p. 117, PI. 20, fig. 6a-e

(here additional synonymy included).

Material. - Over two hundred specimens, partly damaged.Dimensions, description and variation are given in BRoTzEN's (1948) and in POZARYSKA'S

(1965) papers.

Occurrence. - In Poland, common in the Paleocene, i. e. Montian, mainly in PolishLowlands, to be rare in the Pamietowo boring. Common in the Paleocene, i. e. Selandianof Denmark, Sweden , V .S. S. R. (Russian Platform), and in the Paleocene, i. e. Thanetian ofEngland. It occurs in the Paleocene, i. e. Montian, Selandian type beds.

Genus LAMARCKINA BERTHELlN, 1881

Lamarckina rugulosa PLUMMER, 1926

1926. Lamarckina rugulosa PLUMMER, MS; J. A. CUSHMAN, The genus Lamarckina , p, 8, PI. 3, fig. 6a-e .1926. Lamarckina rugulosa PLUMMER ; H . J. PLUMMER, Foraminifera of the Midway , p. 140, PI. 9, fig. 3a-c.1965. Lamarckina rugulosa PLUMMER; B. McGowRAN, Two Paleocene. .. , p. 57, PI. 5, fig. 6.1965. Lamarekina rugulosa PLUMMER; K. POZARYSKA, Foraminifera and biostratigraphy. . ., p, 116, PI. 24. fig. 4a-c

(here additional synonymy included).

Material. - About 40 specimens, usually well preserved.Dimensions, description, variation and remarks are given in POZARYSKA'S paper (1965).

Occurrence. - In Poland, not so common in the Paleocene (Montian only) of PolishLowlands and Parnietowo boring, where it is extremely rare. Known from the Paleocene ofAmerica (U.S.A.), Australia and U.S.S.R. (Georgia). Lamarckina rugulosa PLUMMER wasfound by the present writers also in the Paleocene sediments of Austria. Thi s species is a goodindex species for the Paleocene, Selandian type beds.

94 KRYSTYNA POZAR YSKA & JAN INA SZCZECHU RA

Genus MISSISSIPPINA Hows, 1930

Mississippina binkhorsti (Rsuss, 1862)(PI. xv, figs. 6-10)

1862. Rosalina Binkhorsti ; A. E. R EUSS, Pala ontologische Beitrage.. .. p. 317, PI. 2, fig. 3.?1958. Mississippina binkh orsti (R EUSS); J . HOFKER, Foram inifera from the Cretaceous , XXXVI, p. 101, Text-figs . 1-11 .

?1959. Mississippina binkh orsti (REUSS); J. HOFKcR, Les Fora miniferes du Cretace , p . 376, Te xt-fig. 39.?1960. Mississippina binkh orsti (R EUSS); J. HOFKER, Les Foraminiferes du Cretace , p. 170, Text-fig. 22.

1961. Discorbis binkhorsti (REUSS) ; V. P. VASILENKO, Foraminifery verchnego mela , p. 37, PI. 7. fig. 5.1962. Discopulvinulina binkhorst i (R EUSS, 1862) ; H. HILTERMANN& W . KOCH, Oberkre ide... , (in Leit fossilien ), p. 335.

PI. 50, fig. 17.?1965. Mi ssissippina binckh orst i (REUSS, 1862) ; K. POZARYSKA, Foraminifera and biostrat igra phy. . ., p. 119. PI. 16. fig.I a-b.

1965. Mi ssissippina midwaye nsis trinitatensis CUSHMAN & R ENZ; K. POZARYSKA, Ibid .• p. 119, PI. 17, fig. 2.

Material. - About one hundred specimens well preserved.Dimensions, description and variation are given in POZARYSKA'S paper (1965).

Variation significant; it concerns the size, sha pe and ornamention. Specimens from thebase of Montian (Selandian) are generally much bigger and lower, being less distinctly perfo­rated on dorsal side than those from the true Montian, which are smaller, higher and generallywith a more distinct umbilicus, who se inlets pen etrate the sutures; their umbilicus is sometimesfilled with a small, regular plug (cf. PI. XV , figs. 6-10). It is not excluded that there are twodifferent , probably related species, or one species morphologically differentiated during th eevolution process.

Remarks. - Spec imen s from Pol and included into Miss issippina binkhorsti (REUSS),especially the se from the Selandian bed s, do not differ from specimen figured and describedby REuss (1862) as Rosalina binkhorsti from the Maa strichtian of Holland. They a lso do notdiffer from specimen described by VASILENKO (1961) as Discorbis binkhorsti from the Danianof U .S.S.R. , as well as from those illustrated and described as Discopulvinulina binkhorsti byHILTERMANN and Kocn (1962) from the Maastrichtian and Danian of Germany, and byPOZARYSKA (1965) who described it as M ississippina midwayensis trinitatensis CUSHMAN& RENZfrom the Maastrichtian and Paleocene of Poland. Specimens referred to thi s specie s by HOFKER(1958, 1959, 1960) from the Maastri chtian of Holland, and by POZARYSKA (1965) from the UpperCretaceous and Paleocene of Poland, di ffer ho wever in shape and ornamentation from trueMississippina binkhorsti (REUSS), being much lower and having an ornamented umbilical side;specimens of M. binkhorsti (Rsuss) are never cove red by tubercle s on the umbilical side, wherethey have not any bands of clear shell material parallel to peripheral margin. For thi s rea son,it seems po ssible that the y deviat e, representing another species.

Occurrence. - In Poland no common in uppermost Maastrichtian up to the Montianof Polish Lowlands and Pamietowo boring. Known to be common in the Paleocene of Den­mark, Sweden, Germany, Belgium, Holland and U. S. S. R. , where it occurs also beginningthe Upper Maastrichtian.

Genus STOMATORBINA DORREEN, 1948

Stomatorbina sp .(P I. X, figs. 1-2)

?l958. Epistomina inkermanica SUCKAJA ; E. K. SUCKAJA, Forami nifery verc hnich sloev .. ., p. 205, PI. 3, fig. 2.?l963. Stomatorbina torrei (CUSHMAN & BERMUDEZ) ; J . E. VAN H INTE. Zur Stratigraphie... . p, 116, PI. 18, fig. 1 a-c

(non Lamarckina torrei CuSH. & BERM. ; J. A . CUSHMAN & P. J. BERMUDEZ, 1937, Further .... p . 21 , PI. 2.

fig s. 24-26).

FORAMINIFERA FROM THE PALEOCENE OF POLA ND 95

Longest diameterShort est diameterHeight of test

Material. - Three specimens damaged .Dimensions of two specimens (in mm):

F. IX/2540·160·150·08

F. IX/2550·170·15

Description. - Test trochospiral , biconvex, more con vex on the ventral side, less ondorsal side, almost round, slightly lobulate in outline, periphery broadly rounded, thickened.Chambers triangular, gently inflated , 6 in number in the last whorl, increasing moderately insize as added, coiled in at least two whorls; sutures of dorsal side somewhat curved, raised,limbate, partly obscured, joining the peripheral thickenings; on the ventral side sutures aredepressed, narrow and radial enlarging near the weakly marked central depression. Near theperipheral margin, there are on both sides regular, ovate depressions, parallel to the margin.Surface smooth. Aperture slit-like at the base of the last formed chamber on ventral side ,extending onto the periphery up to the umbilical depression s, which are somewhat asymmetrical.

Variation not known due to the scarcity of material.

Remarks. - Specimens, assigned to Stomatorbina sp. from Poland, are the most similarto tho se included by Suckaja (1958) into Epistomina inkermanica from the Dano-Montianof the Crimea CD. S. S. R.), differing from them, however, by being flatter, not concave ventrallyand ha ving less chambers. In comparison with specimens figured by VAN HINTE (1963),described as Stomatorbina torrei (CUSHMAN & BERMUDEZ), specimens from the Polish Montianare somewhat higher ; in the latters, the inner whorls are not visible.

Occurrence. - In Poland, a few specimens in the Paleocene (Montian) of Pamietowoboring only.

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MARIE. P. 1937. Sur la faune de Forarninif'eres du Calcaire Pi solithique du Bassin de Paris. - Bull . So c. Geol,Francei S, 7, 289-294, Paris .1950. Sur l'evolution de la faune de Foraminiferes des couches de passage du Cretace a u Tertiaire. - Proc.Inst. Geol. Congr., I S, p, 50a , London.1964. Les facies d u Mon tien (France, Belgique, Ho llande). - Mem, Bur. Rech , Geol, Min., 28, 2, 1077-1096(Colloque Paleogene), Par is.(unpubl.) . Foraminiferes d u Calcaire Pisol ith ique (Bassin Paris ien , Belgiq ue, Ho llande). 26 plan ches.

FORAMINIFERA FROM TH E PALEOCENE OF POLAND 99

MARLIERE, R. 1939. Cont ribution a l'etude des form ations cretacees et terti aires du Bassin de Mons. - Ann . Soc. Geol.Belg., 63, 52-78, Liege.1954. Le Cretace (chap. Xll). Le Paleocene (chap. XIII ). - Ex tr. Prodrome Descr. Geol. Belg ., 417-449, Liege.1955. Definition actue lle et gisement du Montien dan s le Bassin de Mons. - Ann. Soc. Geol. Belg. , 78, 297-316,Liege.1957. Sur le "Montien" de Mons et de Ciply. - Bull. Soc. Beige Geol., 66, I, 153-166, Bruxelles.1958. Ostracodes du Montien de Mons et resultats de leur etude. - Mem. Soc. Beige Geol., 5, 1-53, Bru xelles­1961. Le Paleocene et le Cretace superieur de la region de Mons. - 7e Col/. Europ, Micr opaleont. , Guided'excurs.,1-9.1962. Le Monti en de Mons : etat de la question . - Col/. Paleogene, 1-12, Bord eaux.

MASLAKOYA, N. 1. 1959. Strat igrafieeskaja schema verchnemelovych otl ozenij Kryma. - Vestn. Mosk. Univ., 1, 109-112,Moskva.

MATWIEJEWOWNA, L. 1934. Anal iza fauny mallow i slimakow siwaka z oklic Pulaw . Strat igraphi sche Betrachtung derPelecypoden- und Gastropodenfauna des "Siwak" in der Umgegend von Pula wy bei Lublin . - Prace Tow. Przyj.Nauk, 9, 91-117, Wilno , 1935.

MCGOWRAN, B. 1965. Two Paleocene foraminiferal faunas from the Wangerrip Group, Pebble Point Coastal section,Western Victoria. - Proc. Roy , Soc. Victoria, 79, 1, 1-74.

- 1966. Austral ian Paleocene Lamarckina and Ceratobulimina , with a discussion of Cerobertina, Pseudobulimina,and the family Robertinid ae. - Contr. Cush. Found. Foram. Res., 17, 3, 77-103, Bridgewater.

Mensa , M. 1959. Sur la limite superieure de l'etage Maastrichtien dans la region-t ype. - Acad. Roy . Belg., Ser . 5, 45,3, 316-338, Bruxelles.

MEUNIER, S. 1912. Geologic des envi ron s de Pari s. 1-540, Par is.MJATLUK, E. V. 1953. Spirillinidy, Rotalinid y, Epistominid y i Asterigerinidy. - Isk op. Foram. SSSR , 71, 1-261, Le­

ningrad-Moskva.MOROZOVA, V. G. 1939. K strati grafii verchnego mela i paleogena Embenskoj oblasti po faun e foram inifer. - Bjul.

Mosk. Obsc. Isp. Prir., Otd. geo!., 17, 4{5, 59-86, Moskva.1946. The boundary between Cretaceous and Tertiary deposit s in the light of the study of foraminifers.­C. R. ( Dokl.) Acad. Sci . VRSS., 54, 2, 153-155, Moskva.1958. K sistematike i morfol ogii paleogenovych predstavitelej nad semejstva Globigerinidea. - Vopr. Mikropaleont.,2, 22-52, Moskva.1959. Stratigrafija dat sko-montskich otlozenij Kryma po foraminiferam. - Dokl. Akad. Nauk SSSR., 124, 5,1113-1116, Moskva.1961. Datsko-montskie planktonnye foram inifery juga SSSR. - Paleont. Z UI"II. Akad. Nauk SSSR. , 2, 8-19,Moskva.

- , KREIDENKOY, G. P. & D AYlDZON, R. M. 1965. Biostratigrafia paleocenovych otlozenij Tad zikskoj depresii.­Biul. Mosk. ObS(~. Isp. Prir., Otd. geol. , 3, 34-55, Moskva.

MOSKYIN, M. M. 1959. Atla s verchnemelovoj fauny severnogo Kavkaza i Kryrna, - Tr. VNllGAZ, 1-304, Moskva.& NAJDIN, D. P. 1959. Stratigraphie du Cretace superieur de la Plate-forme Russe de la Crimee et du Caucasedu Nord. - 84e Congr, Soc. Sav. , 497-522, Dijon.& - 1960. Datskie i pogranicnye s nimi otlo zenija Kryma, Kavkaza, Zakaspijskoj oblast i i jugo vostocnojcasti Russkoj platformy. - Meid. Geol. Kongr., 21 Sess ., Dokl. SOl'. geol., 15-40, Kop enhaga.

NAGGAR, Z. R. EL-. 1966. Stratigraph y and planctonic Fora minifera of the Upper Cretaceous - Lower Tertiarysuccession in the Esna-Idfu Region, Nile Valley, Egypt, U. A. R. - Bull . Brit . Mus. (Nat, Hisr.) , Geology,Supp!. 2, 1-291, London.

NAKKADY, S. E. 1957. Biostratigraph y and inter-reg ional correlation of the Upper Senonian and Lower Paleocene ofEgypt. - J. Paleont ., 31, 2, 428-447, Mena sha.

- 1958. Stratigraphic and Petroleum Geology of Egypt (U.A. R.). - Univ, Assiut, Monogr . ser., 1, 1-215, Assiut.- 1959. Biostratigraph y of the Urn Elghana yern section, Egypt. - Micropaleontology , 5, 4, 453-465, New York.

OLSSON, R. K. 1960. Foraminifera of latest Cretaceous and earliest Tertiary age in the New Jersey Coastal Plain . ­J. Paleont., 34, 1, 1·59, Menasha.

ORBIGNY, A. D'. 1846. Foraminiferes fossiles du bassin tertia ire de Vienne. 1-152, Paris.PLUMMER, F . B. 1932. Ceno zoic systems in Texas. The geology of Texas. - Bull. Texas Univ., 3232,

Austin.PLUMMER, H. J. 1926. Fo raminifera of the Midway Formation in Texas. - Ibidem, 2644, 1-204.

1934. Epistominoides and Coleites, new genera of Foraminifera . - Amer. Midi. Natur ., 15, 5, 601-608, Lancaster.- 1936. Structure of Ceratobulimina . - Ibidem, 17, 2, 460-463.

7*

100 KR YSTYNA POURYSKA & JANI NA SZCZECHURA

POZARYSKA, K. 1952. Zagadnienia sedymentologiczne g6rne go mastrychtu i danu okolic Pulaw (The sedimentologica lproblems of Upper Maestrichtian and Danian of the Pulawy environment , Middle Vistul a) . - Biul. P. Inst .Geol. , 81, 1-104, War szawa.1957. Lagenidae du Cretace superieur de Pologne (Lagenid ae z kredy g6rnej Polski). - Palaeont . Pol., 8, 1-19C,Warszawa.1964. On some Foraminifera from Boryszew boring, central Poland (0 pewnych otwornicac h z wiercenia w Bory­szewie). - A cta Pala eont . Pol., 9, 4, 539-548, Warszawa.1965. Foram inifera and biostrat igraph y of the Danian and Monti an in Poland (Otwo rn ice i biostr atygrafia danui montu Polski). - Palaeont . Pol., 14, 1-154, Warszawa.1966. The Cretaceous-Tertiary bound ary in Polan d and adjacent areas (in the light of Foraminifera). ­A nn. Soc. Geol. Belg. , 89, Bull. 3, 47-64, Liege.

POZARYSKI, W. 1938. Stra tygrafia senonu w przelomie Wisly miedzy Rachow em i Pulawam i (Senonstra tigraphie imDurchbru ch der Weichsel zwischen Rach6 w und Pulawy in Mitt elpolen). - Biul . P. Inst . Geol., 6, 1-94, War­szawa.1957. Poludniowo-zachodnia krawedz Fennosarmacji. The south-western margin of Fenno-Sarmatia. - Kwart .Geol., 1, 3/4, 383-424, Warszawa .& POZARYSKA, K. 1959. Comp araison entre le Cretace de la Belgique et de la Pologne. - Ann . Soc. Geol, Belg. ,82, 1-14, Liege.& - 1960. On the Danian and Lower Paleocene sediments in Poland . - Int , Geol. Congr., 21 Sess ., Norden,5, 170-180, Cop enhagen.

RASMUSSEN, L. B. 1960. Geology of north-eastern Jylland , Denmark. - Ibidem, 2, Guide excurs. A42, C37, 1-38.RAUZER-CER NOUSOVA, D. M.& FURSENKO, A. V. 1959. Prostejsie. In: J. A. ORLOV (red.), Osnovy Paleon tologii. 1-482,

Moskva.RAVN, J . P. J. 1925. Sur le placement geologique du Danien. - Publ. S erv, Geol. Danemark , Ser. 2, 43, 5, 1-48, Kjo­

benh avn.REICHEL, M. 1952. Remarques sur les Globigerines du Danien de Faxe (Danemark) et sur celles des couches de passage

du Cretace au Tertiaire dans la Scaglia de I'Apennin. - Eel. geol. Helv., 45, 2, 341-349, Zurich 1953.REISS, Z. 1960. Structu re of so-called Eponides and some other Rotaliiform Foraminifera. - Geol. Surv. Israel, Paleont .

Div., Bull. 29, 1-24, Jeru salem .REUSS, A. E. 1861. Palaontologische Beitriige. 1-3. - Sitzber. k , Akad. Wiss., 44, 304-342, Wien ,ROSENKRANTZ, A. 1920. Craniakalk fra Kob enh avns Sydhavn. - Danm . Geol. Unders., 2, 36, K obenhavn.

1924. Nye lagttagelser over Cerithiumkalken i Stevns Kl int med Bemaerkninger om Graensen mellem Kridt ogTert iaer. - M edd. Dansk. Geol. For., 6, 28-31, Kebenh avn.1937. Bemaerkninger om det ostsjaellandske dan iens stratigrapfi og tektoni k. - Ibidem, 9, 2, 199-212.1939. Faunaen i cerithiumkalken og det haerdnede skrivekridt i Stevns Kl int. - Ibidem, 9, 4, 509-514.1960. Danian Mollusca from Denmark. - Int . Geol. Congr. , 21 Sess., Norden, 5, 193-198, Copenhagen.1964. Note on some Cranias from central Poland (0 pewnych kran iach z Polski srodkowej). - Acta Palaeont .Pol., 9, 4, 513-531, Warszawa .

ROUVILLOIS, A. 1960. Le Thanetien du Bassin de Pari s (Etude hydrogeologique et micropaleontologique). - Mem .M us. Na t, Hist., N. S., 8, 1-151, Par is.

SCHMID, E. M. 1962. Die Foram iniferenfaun a des Bruderndorfer Feinsand es (Danien) von Haidhof bei Ernstbrunn,NO. - Sitzber, Osterr. Akad. Wiss. , Matli-Nat. KI., Abt. I, 171, 8/10, 315-361, Wien.

SIGAL, J . 1958. Reflexions it prop os des termes Paleocene et Dan ien. - C. R. Soc. Geol. France, 5, 94-97, Pari s.STAESCHE, K. & HILTERMANN, H. 1940. Mikrofaunen aus dem Tert iar Nord westdeutschlands . - Ab h, Reichsst , Bodenf. ,

N. F., 201, 1-26, Berl in.SUII UOTlNA, N. N. 1947. Foraminifery dats kich i paleogenovych ot lozenij scvernogo Kavkaza. - Mikrofauna SSS R,

Tr. VN IGRI, 39-160, Leningra d-Moskva .1950. Mikr ofaun a i stratigrafija Elburganskogo gorizon ta i gor izont a Go rjacego kluca. - Ibidem, 4, 51, 1-109.1953a. Verchne-eocenovye lagenidy i buliminidy juga SSSR. - Ibidem, 6, 11 5-255.1953b . Glob igerinidy, Hantkeninidy i Globorot ali ty. - Iskop, Forum , SS SR , Tr. VNIGRI, N. S., 76, 1-239,Lcning rad-M oskva.

SUCKAJA, E. K. 1956. Stratigrafija nizn ich gorizontov paleogena centralnogo Predkavkaz'ja po foramin iferam . - Tr.GIN AN SSSR , 164, Moskva .1958. Foram inifery verchnich sloev "datsko-mo nskich" izvestnjakov jugo-zapadnogo Kryma . - Tr. VNIGNI,9, 197-210, Moskva.1960. Fora minifery verchnego paleocena jugo-zapadnogo Kryma (Bacheisarajskij ra ion). - Ibidem, 16, 235-259.

FORAMINIFERA FROM THE PALEOCENE OF POLAND 101

SZCZECHURA, J. 1965. Cytheracea (Ost racoda) of the upperm ost Cretaceou s and lowermost Tert iary from Central andNorth Pol and (Cythera cea (Ostra coda) najwyi:szej kred y i najnizszego trzeciorzedu Pol ski srodkowej i p6!nocnej).­Acta Palaeont , Pol., 10, 4, 451-564, Warszawa.

TERQUEM, 0 .1 882. Les Foram iniferes de l'Eocene des environs de Pari s. - Mem. Soc. Geol. France, Sec. 3, 2, 1-193, Par is.TOULMIN, D . L. 1941. Eocene smaller Foram inifera from the Salt Moun tain limestone of Alab ama. - J. Paleont., 15,

6, 567-611, Menasha.TZAGAREU, A. L. 1954. Verchnij mel Gruzii. - Akad. Nauk Gruz: SSR, Monogr., 5, 1-462, Tb ilisi.TRo ELsEN, J . C. 1957. Some planctonic Foraminifera of the type Dan ian and their stratigraphic impor tance. In: LOE­

BUCH, A. R. , TAPPAN, H., BECKMANN, J . P . et al., Studies in Foram inifera. - U.S. Nat, Mus. Bull., 215, 125-132,Washington.

VASILENKO, V. P. 1950. Foram inifery paleocena centralnoj cas t i dnep rovsko-doneckoj vpadiny. - Mikrofauna SSSR,Tr. VNIGRI, 4, 177-224, Leningrad.

- 1954. Anomalinidy. - Iskop. Foram. SSSR, 80, 1-203, Moskva-Leningrad.- 1961. Foraminifery verchnego mela poluostrova Mangyslaka , - Trudy VNIGRI, 171, 1-487, Leningrad.

VINCENT, E . 1930. Etudes sur les Mollusques montiens du poudingue et du tuffeau de Ciply, - Mem, Mus. Roy .Hist. Nat. Belg.• 46, Bruxelles.

VISSER, A. M. 1951. Monograph of the Foraminifera of the type local ity of the Maestrichtian (Sout h-Limburg, Ne­therland). - Leidse Geol. Med., 16, 197-359, Leiden.

VOlGT, E. 1960. Zur Frage der stratigraphischen Selbstandigkeit der Danienstufe. - Int. Geol. Congr., 21 Sess., Norden,5, 199-209, Copenhagen.

- 1964. A Bryozoan fauna of Dano-Montian age from the boring Boryszew near Sochaczew in Central Poland.­Acta Palaeont , Pol., 9, 4, 419-498, Warszawa.

VOLOSINOVA, N . A. & DAIN, L. G . 1952. Nonionidy, Kassidulinidy i Chilostomellidy. - Iskop. Foram. SSSR, N. S.63, 1-151, Leningrad-Moskva .

WIENBERG RAsmISSEN, H. 1962. The Danian affinities of the Tuffeau de Ciply in Belgium and the post-Maastr icht ian"Me" in the Netherlands. - Coli. Paleogene, 1-8, Bordeaux.

WiLUAMSON, W. C. 1858. On the recent Foraminifera of Great Britain. Ro y, Soc. of London, 1-99, London.WOOD, A. & HAYNES, J . 1957. Certain smaller British Paleocene Foraminifera. - Contr. Cush, Found. Foram, Res.,

8, 2, 45-53, Washington.

ALPHABETICAL INDICES

r. INDEX OF AUTHORS

Page

C

CAl.VEZ. Y.I.E. . 49. 50. 61, 62. 72. 75, 82CHALlLO\'. D. M. . 30CORNET• .I. . , , . . . . . . . . . 26. 28CUSHMAN• .I. A. 22. 23, 3'5 . 3li . 39, 44. 45. 48. 50,5 1. 55.

56. 58. 59. 80. 82. 83. 85. 93. 94Cl;VILLl ER . .I. . . . . . . . . . . . . . fi8 . 27. 33. :\5. 55, 64, 72

HANNA, C. D . . . . . . 85HARDER, P. . . . . . . 28HAYNES, J . 17,1 9,34,44,45,47.48.52,53,76, 81, 87HEINZELIN, J. DE . . 26HERMANNI. E. V. . . 85HILLEBRANDT, A. v. 17HILTERMANN, 1-1 . . . 27. 94HINTE, J. E. VA N 94, 95HOFKER, J. 16. 17, 26-31. 33, 34, 36. 37. 39, 42, 45, 49-55.

58-60, 62. 63. 65. 6fi. 69. 70. 72. 78. XO. X6. 87, 89,90, 91, 94

I-I USEZIMA. R. . . . . . . . . . . . 5 1

Pa ge

F

FORCHHAMMER, G . 28FORNASINI, C. R. 62, 63, 66FRANKE, A 16

G

G LAESSNER, M . F. 88GLIBERT, M . 26GLINiZBOECKEL, C. 68GRIGELIS, A . A. 20GROSSOUVRE, A . 28

H

I ;o.l I>AI'S. .I.

. 24

A

B

BRONNIMAN:". P. . . .BRUNNICH NIELSEN. K.BYKOVA. N . K.

ABR ARD. M. R. . . . .

BANG, 1. .BATJES, D. A. J .BELLEN, R. C. VAN 10, 26, 27,

... .. .. 16, 2X

. . . . . .. . . 7636, 37, 50, 51, 54-56,

60-67, 72, 74, 80BERGGREN, W. A. 28, 33, 34. 69BERMUDEZ, P. J . . 55. 56, 58, 59, 68, 94BEZRUKOV, P. L. . . . . ... 19BOLLI, H . M . . . 21, 22, 66, 68BRIART, A. . . . . . . . . . 28BROECK, E. VAN DER . . . . . . . 31BROTZEN, F.9, 10, 14, 16, 19,22-24,26,28,29, 33. 34 37,

38, 40, 44-49, 52-54, 58. 64, 70, 72, 77-80, 83-87,89-91. 93

6928

22. 30

79. 92212120309422

D,\LIlI I'/ . F..DAM. A. TENDA~10TTE. R .

D,\VIDZO:". R . M.

D r.uoo, G .DESOIl. E. . . . .

EIlWARI>S. P. G .EGG ER. .I. G .ELI .IS. B. F. . .

1)

E

. . .. .. . 6827. 5 1. M. 75. 79. 81. 82

24

14. 26, 30. 3 128

. . . . . . . . 48

. . . . . .. . 4241-43. 46. 52, 60-63. 82

.1

.IANSI I' . A. L. . 34

.IEN:"I:"GS. P. H. 36. Xfi

K

K""SSCHI ET EIt . .I. 1'. 1-1 . 49. 50. 66. 72-75. 77.K A~ARAVA , I . . . . .KA~ARAvA , M.KAPTARE;o.lKO-C ER ;o.lOUSOVA. O. K.KELLER, B. M . . . .KOCH, W .KREJ DENKO\' . G . 1'.

FO RA M I N IFE RA F R O M TH E P ALEOC E N E OF POLA ND

Page

103

Page

R

R ASMUSSEN, L. B. 16, 28R AVN, J. P . J . . . 28R EISS, Z. . . . 72, 73R EUSS, A . E . 35, 59, 94R OEMER, F . A . . . . . . 45R OSENKRANTZ , A . 10, 16, 17, 26, 28-30, 33, 34R OUVILLOIS, A . 33, 41, 43, 50, 51, 72, 74-78, 81, 92R UTOT , A . 31R Z EHAK, A . . . . . . . . . . . . 91

LMI,\R CK, J . B.

L EMOINE, P. .

Lroxov, G . P. .L ERICHr . M . . .

LOI-Il U CH, A. R.

M ARIE, P . .. 24, 27,MARLIERE, R.MARSSON, T.

M ARUH ASI, M . .

MASLAKOVA, N. I.MCGOWRAN, B.M EIJER, M • . .

M ESSINA, R . A .

M EUNI ER, S . . .

M OROZOVA, V. G .M OSKVIN, M. M.

L

62, 6324192651

i\l

30, 36, 37, 41, 54-57, 62, 63. 10, 14, 26, 30

395120

23,47, 51, 76, 85, 87,91, 93. . . 31

41-43, 46, 52, 60-63, 82.. .. . 2420-22, 30, 37. . .. . 30

N

S CHMJD , M . E.S COTT, G .. .

SMOUT, A. H . .

STAESCHE, K . .S UBBOTl NA, N . N.S UC KAJA, E . K.S ZCZE CHURA, J . .

s17,49, 67-70, 93

· . 2262, 63

· . 2768, 71

20, 32, 89, 90, 94. 95. . . . .. . . . 30

oOLSSON, K. R.. . . . . 40, 68ORBIGNY, A. D' . 41-43, 46, 47,52,53,60-64,72, 73,76,82

p

P HlLIPPI , R. A. . . . . . . . . 37PLUMMER, F. B. 22hUMMER, H . J. 22, 39,40, 68, 70, 71, 83, 89-93PONTON, G. M . . 35POSLAVSKAJA, G. G. . 30POZ ARYSKA, K. 9, 10, 14, 16, 17, 2 I , 1.6, 29-3I, 33.

35-41,43-48,50,59,60, 69-74,76,79, 81, 83·91,93,94POZ ARYSKI , W . 29, 33

WIENBERG-RASMUSSEN, H . .

WILLlAMSON, R . . . . . . .

NAGG AR, Z. R . EL-.

NAJDIN, D . P. . . .

N AKKADY, S. E. . .

NIELSEN-BRVNNICH, K.

69-71303328

T APPA N, H • ..

T ERQUEM, O.

TOULMIN, D. L.TROELSEN, J. C.

TZAGARELI, A. L.

VASILENKO, V . K.VASILENKO , V. P .

VINCENT, E •.

VISSER , A . M . . .

T

. . . . . . . . . . 5149, 50, 57, 58, 61, 62, 75, 81, 82

· . 8428, 68

· . 21

v. 19

19, 20, 34, 77, 88, 89, 9426, 30

. 50, 51

w26, 30, 31

57

11. PALAEONTOL OGICAL IND EX

Systemat ic names c ited oni ~ ale indica ted by italic. and na mes accompa nied by des crip tion s arc indicatedby roman lette rs.

Numerals which indi ca te the page. on which the species is cited on ly, ale sta ndard ; nu mer als, which indicat epages with descriptions, a rc bold.

Numera ls den oted with aster isks ( *) indica te pages with figures.

Page Pa ge

c

1" -22. 27, H6. 8H, PI. X I\'868686

22 . 25, 6939

2 1,22,25 , 39

acuta , A nomatinaucuta , Globorotulla ,acu t ldorsatum • No nionacutus, Eponides ,Atabanuna . . . .amygduloidcs , Globu tina

(Jug ula/ a , Globo rotalia .Angulogerinu

an gusta.• Pseudop o lvm orph ina gcii er iAnomalina .uppl inae, LIJXO~/Olllf{m

arcuata, M nrginulin aa rcua tus , Astuco lusarenacea , G lobulinaA renobuliminaark adelphkm a , Bulimi"0a rka nsasana, Tcxtulan a plum mera carmata, Mo rs inutina er.Astacolus . . . . .as tero ides , Ci bicidesuudouini , Rota/fa

uurouzae , C ibici desauroueae, Clbicklcs (Cibicidina )

Bbeanmonsunu , Cibicidesbiforrni s, Frondicula ti u .bigibbe ra, Alubamlna.binckhors ti • •'/ j.\ ,\ is~ iJ'fJ jJ1a .

binkhorsti, Discaputvinu lma

bink ltors ti, Disco rb is

binkhorsti , 1\1ississippina .

binkhorsti , Rosul inu

BotivinaIJu/ir illo ide.\

breslesens is, Cib icidcsbrorre ni , Conorb inabro tzeni , Rosal inabrotzcn i , Sigmomorphi na

Bulnnina .bundcnsis, Discorbiv . .bundensis, Epistomuri a .

bundcnsis , ROW/ill . . .

bnndcns is , Rotorhinelta .burli ngtoncusis, Anoma lina .

burllngtoncnsiv, Ci bic ide s

ss()~

S250 . 5 1

H4

2017, 21. 22 . 6S

47 , 4817, 44

86 . 8817. 21. 22, 25

37

3717, 4U

3617

23 , 353937

25 . 73, 74, PI. Xl. .. . . 6625, 74 , P I. X

. . 74

"23. 39729494

94

94 . PI. XV

9425, 47

25n . n

5 151. 52, PI. 11

17. 45. 47

.. 5525 . 74 *

· . . 54· 54, 55

19. 86, 87. PI. X VI

· 22. H6

cant ii, Cibicides .. .cari nata, Trnn comlinaca rina tus, Cibicides .cusint irovens is, Belemnella ,cassivellauni . Cih icidesCeratobu litni na . .Cerithiumchoct avens is ornata, C ilncides

CibicidesCiblcides sp ,

Coleit es

cO/w num'. Non ion .co mmunis comm unis. No nionellaco mmunis. G uuulinacannnunis , N O/lion . .

co mmunis pa lcocenica , Non ion ellacompressa. Globigerlna , . . . . .cornpressa, Gl o bigc rina aIT.compressa , Globorotalia ( Turborotalia )conico truncata , Globorotulia . . . . .constri ctus vulgaris, Hoptoscaphi te ,co rd ieriana na varr o unu, Osan gul.u iuco r, ugata , An otnalinucorrugata , Disco rbis . .corrugata, Patet ltnaco rrugnta. Rotorblnolt a .costa ta , Anomulina um bilica tacostulara, M argjnulin a . . . .c rassa , Pyrami di narryntoniphalus cl'. berceunvinensiv, Cibictdc:\cubensis , Boldtucunea ta, AIlgu hH,:erinu

cuneata , Pyra rnid jna

cunobelini, Cibtcidc- :c usk leyae, Areno bu tim innc uvillieri , Ctblci ucs

Cm'ill ia illu.

D

da nica , An orn ali nadanic a. An omalino idesdanica, Cibicides . .danlca, Gavelinetta .da ub jergens is, Gl obigerin adego ly eri, Cristellar iadeg o 'yeri , Robulus

. . 25, 7675

1'1. 23, 75 *, P I. X I29259.12S747.1

73, 769082

H2, 83 , PI. IX42*. 4 .1

8383, PI. IX... . 6S

11, 25,6H'692 129

11. 1(" 17,1 9.21,22,85

· 55, 5627 . 55. 56

575:'

89 , PI. X III, ')

17, 19, 4676, 74

. ... 2517, 46. 47

· 17, 46.. 25

· 16 . 36 *25. 76, 77. PI. X II

5S . 59

FORAMIN IFE RA FROM THE PALEOCENE OF POLAND 105

Pa ~c

1\1

.. . , ... 36 , 3725·27 . 36 . 37, PI. II I

· . . 16. 17

· . . .. 6811. 16 , 19. 52 . PI, I1

· 11,25 ,70 '

464620

63 . 64 . PI. IX63. 64

. . . 93"0 . 21. 25, 77 , P I. X

· . 7711, 16, 25 , 9 1. PI. XII

SO2.1. 811 . PI. X I

1'1 "I. 2.1. 25. 511 *, 5 15076

20 , 22

1794

S9, 90515 1

26 87" . SS. PI. XI\9';

11 zs.», 55 56, 1'1. VIS(), 57

41. 1'1. I II4 1

644'1

5S. 5~

SS. 5'111. 24·26 (d . Id hh. 67 PI>. V, XVII I

393'1S7

D ,8754 , 5554 . 55

: 5 . 26. 54 , 55 , l' ls . IV, XVII

· . 74· . 73

11. 17. 1'1·23. 84 '20, 22, S9, 90

L

k eiizeri, M urssonella , .keijzer i, Vacuovalvulinuklagsha mn ensis , R obulu.\

k ochi, Globorotalia . . .koeneni , R osal ina . . .kozl owsk ii, Gl obigerina

Iaevigat n , Glandulina .Iaevigata , Xo doso ria (Gtandntina )

laevis , Sp iroplec tunnninu vuriuta cl'.larn arck i. Elph id ium e t,knnurck it . Xonionina

Lamarcl- Lno , . ..

lectus, C ibicides . .teems , Cihicides ( Cib icidoide.\ )

lc llingcnsis, Guvelincl lu .

timbutu, Guvclinello

lim bat a , Pla nulinalim burgensis. Episto r nincl!a e t.limb urgcns is , Pseud oparrrllo ,

Iobatulus, Cibicidcstuna tu.... Cibicides .

nutdruguensis ; Botdia

mad rugaensis, Rota/ill

ma drugaensis , Th alma nnitu

mur gina ta , Ror al iaM urglnul ina .M aru inut inc sp. B .

ma riue, Cib icidesma rtae , Cib icldinamu riei , Parrell a .

nuirtei , Po zarysk aio

rnar ie i. Rot orbinclla .mauricensis, Cibicidesmauricens is , Cibicides sp. cl.rnidw ayensis , Alabam inamidwayensis, Anomal inamidwayensis, Bullmina ,midwaycnsis trlntratensis , Af is" i.,sipfJllI ,1

snidwaye nsi s , Truncatntina,

mi nima , Epollides

minirnns, Ep onides .

mi nor , Anom al inu .

M ississipp inu . .rno ut iuna, Ro to rb incll ununuis , Rotorbinoidv»m ulusu ia ra, Globul ina gibbnmn lt istr iata , Gtobnlinu gihl", V..i L

mult ivuturu tum • Xo nion .mnralis . A l1gulogerillo

17 20. 0 ' 91*. 92_ 0 .

2 14444

17. 443'1

17. 21. 25. 45

9 120

17, 4464

4120. 41

41. 1'1. III4142466869

25 2;' , 65 . "I , VII65

6566 , PI. VI

67, 6867 , 68

254U

42 1',. III8 1

11. 1'1. 25 . 81* . 1'1. IX86

11 25 67 68 . 1'1. VII I72

27, 72

27 . 8616 . 20, 37

4285

. 57

. 4916 , 20 . 21. 23 , 25

17, 19. 22. 23, 87 , PI. XI II87876463

50 , 51. . 72

11.1 7.1 '1. 23.48.49,1' 1. I

. 17, 254'1

E

F

G

dentata, Pa tell ina

Discorbi, . . . .d iscus , Plu nulano

ekblorn i, Anom alinaekbloini , Cibicidcs .ekblomi, Govctinetla

Elphidietlu .Elphidi unt

Epistominellu

Eponi des .eu ro paca , Angulogcrlnacuropaea, Pyramidino(' lI r opoeu. Reussclta .

Ga velinella . .geijcr} , D tof ro ndicuta nu

geijer i a ngusta , Pscudo po tymorphiuagel eenense , Nonion , . .e tbba , Glabulina . . . . .gibba gibba , Gl obulinagibb a multi st riat a . G lobu luu. . .gibba va r. multist riu tu , Gtobntina

glbba vac. striata , Gtobutinu

Glandulina . .Globig erinu . . .Gtob.e erina s p .

glo bigcrinifo rm is . Gl oborot.rlu,globigerinifo nnis , Pararotutu,

Globorota liaGl oborotalia SI'.

Globorotalite s . .Globo rota ti tcs ? n. sp.Gtobo tmncuna

Gtobu lina ,

G lob ulin" sp.uranife ra, N onioninagra n lfcrum , Noni ongr(Jnosa . Anomulinu

grun u lar us, Gt oborotulue,

grat a . Rotalia . . . .gra tu s , Epo nidess ross erugosa , An om ah na

g l yi, Astacolu sGuttu lina . .Gvr otd inoides

fall ax, Karrcr ia .

fr ankei , Enonidesfrond ea , Boli vina,

trondea , PolymorphinoIrondea, Pscud op o lyrnorphiua .

From/feu/ar ia . .

tu sifo rmis , Pyrul ina .

11huil i , A llomorphinu ..

hercegovinensis , Cibicides cry plump!ltllu.' ..:1.

incrassata . Bolivi nutntccn nanica . Epist untinu

Karr erla , . . . . . .kc ljzeri, Conicovalvulina

17. 2277

9. 11'14,95

9136, 37

nahrolensis , La marck ina

nnva rroa na , O san gula ria co rd lerian u 11fl(1\·arroarza, Pul vinulincl lu .

N eo eponides

Neoflabellina . . . .nob ilis , A nomatinoidcsnod ifera , Spiri llina c f.

nodif eru , S pirittina .

N on ion . . . .Nonionclla . .Non ionellu sp .

16 , 22. :!S16 17, 1' /. " I, 22, 85

85722519

11.58. PI. III5S8 18283

106

o

KRYSTYNA POZ AR YSKA & JA NIN A SZCZECHURA

Pag e

R

Page

octocamerata I Gy roidinotdes ,oedurni, Boli vina . .orbicularis, Cibicides .ornata , Cibicides cho ctavens is

ornatus , CibicidesOsa ngularla , . .ova ta , Bulimin aova ta, Nonio nellaorata , Praeglobuliminuodtuni , Bolivlna . . .

p

16, 22, 8517, 19, 23,47

75747485

17, 19, 2 1, 22 , 47 - , 4817, 83

47

· . 47

ranco casensis, Robulu srcticulosa , Pul vinulinaret iculosus , Coleit esRobu/usrobusta , Pal mula ,Rosalina . .

R omlla . . . .Ro torb inella . .Rugoglobigerinarug osa, Gaud ry lna

rllgosa, Rugo globigerina

rugulosa , Larnarckin a .

40­90

17, 23, 27, 9014, 39

· . 1651 , 53

60542520

. 9, 1111, 14, 21-23, 25 , 93

tegulatus; lnoceram usTextularla

thouini, Rota lia . .Thuhnanni tatorrci, Lamarckin asorrel, S tomo torbtnatoulmini, Epo nides 16. 17,triloculinoides, Globigerinatriloculinoides, Globigerina (Subb otin a)t rinitatensis , Discopul vtnulina . . . . .t riniunensis , Mlssissippina ntidwayensis ,t rivial is, G lo bigerinaTrochann nina sp. . . . . .t roc h idifo r rnis, Rotaliatubercul at a , Ceratobul iminatuberculata , Globulina . .tu berculat e , G lo buli na cf.tubcrculifera, Pararotaliatuberculifera , Rotali a .turbinat a, Criste llariaturbi natus, Robulusturgida, No nionlna .Ty locidaris . . . .

paleoceu ica , Bulimina . . . .

paleocenica , Bulimina ( Reussel/a ) .paleocen ica, Pseudopol yrnorphina .pale ocenica , ReusselJa . . . . . .

paleoceni ca , N on ionella co mmunispale ocen icus, AstacolusPa/mu/a .papil lat a , Rotorbinella .Pa rarotal la . . . • . .porisicnsis , Neoconorbina tRosallna}parisiensis, Rosalina

Patellina .Patellina sp. . .pegwellensis , Hollandinaperovalis, Rotalia

plana I Discorb is .Planu taria ,Planulina . . . .plummerae, Tex tulariaplummerae arka nsasa ua , Te xtulariaplumoides , Citharina : . . •porneraniana, Cuvillierina? .

pommerallu. St ensiolna .po nto ni , Gyroidinoides.Poiaryskaiapraeacuta, Anomalina .praepseud ornenardii, Globor otaliaprima, Siphonina (Pulslphonina} cf.prima, Elphidiella . .prim a, Pulsiph on ina .prima , Sip boninu

primum, Elphi diumpr oblerna , Guttulin a .propr ius, Cibicidc sprop rius , Cibic ides ( Cibicidoides)pseud obullo ides, Gl o bigerin apseudobullotdes , Globorotalta

pseudodiscoides , Discorbis . .pse udodtscoides , Gaveiinops ispseudo -mutnilligerus ; Robulus

pseudomenardii, GloborotatiaPseudoparrella , . . . . . .Pseudop olymorphina . . . .pseudoregularis, Sigmomorphinapulavensis pulav ensis , Planularia .Pul siphonina ,PyramidinaPyru lina ..

Q

q ua drat a , D iscorbisquadrata , Rosalina

17, 19, 20, 22 , 25, 4848

17, 25, 44. ... 2083 , PI. IX

11, 16,38 -14

11, 25, 26, 56, 57, PI. VI59

52, 5352, 53

5711, 57, PI. III

· . 2561,62

80

· . 14· . 8027,3523 ,3522, 23

11, 25, 26, 58 , 59, PI. IV. 9, 11

· . . 16,85· . . . . 54

17, 19-21 , 23, 8811, 25, 26 , 66 , PI. VI. . . . . . . 92, 93

17, 23 , 25 , 27, 64, PI. IX11,1 7,19,20,22,23,92

. 92

. 6443 -

17, 19, 20, 22, 77, 78, PI. XVI. . . . . . 77

17, 22, 25 , 70· . 7060,61· . 60

17, 21 , 22 , 2322, 66

5144

17, 25, 4516, 25· . 9246, 47

· . 45

49, 50, PI. II· . . . . 49

sa hlstr oern i, Cibicidessahls tronu, Clb icides .saxo rum. Rot alia . .scalaris, Hoeglundinaschreiber si, Eponides .schreibersi, Neoeponidesschreibersi, N coepon ides cf.schreibersi, Rotalina .sela ndiana , R osalinaselmensis , Tappanina .Si gm omo rphina . .simplex , An omalina .simplex , Cibicide s . .solutu , Sigmomo rplrinasoldanii , GyroidinaS pirillina . . . .S plroplectanunina

S tensioina . . . .St oma torbina . .Stom ator bina sp.striata , Globulina gibba var ,sub involutus , Cibicides .su blaeve, Nonion . . .sublaeve, Prote lphidiu msuccedens, Cibicides . .

s

T

. 17, 19, 23 , 78 ­. . . 78

25-27, 61, 62, Pis. V, XVIII17, 21-23

· . 7272, 73

72, PI. XV· . 72, 73

19, 25 , 52 *, 53, PI. II17, 19,23

. . . . .. 45

. .. . 89, 9017, 22, 23 , 78

172058352594

94, 95, PI, X· 42

. 7611, 81, 82 , PI. IX.. . . . 25,81

17, 19-22, 25, 79 -

2935

62, 63

· . 59· . 9494, 95

19, 23 , 25 , 27, 72, PI. XV11, 17, 19, 22, 25 , 71

71

· . 80· . 94

11, 22 , 7I· . . . 50

24-27, 62, 63 , PIs. V, XVII11, 14, 17, 19-21, 25, 93

. 42

. . . . 42, PI. III25-27, 59- , 1'1. VII

59, 60

· . 4023,40

83

· . 29

FORAMIN IFERA FROM THE PAL EOCEN E OF POLAND

Pa ge

107

Page

uubiquu , LJiscorbis . . . . . .

umbilicata, Cibtctdes : . . . .umb ilicat a cos ta ta, An ornal inaumbilicata, Gavelinella . . . .umbiIicata umbilic ata, An omalina .

vVacuovalvulinaVagin ulina sp .vallensis , Laff i tcina

. 50

. 8989, PI. XIII

. . . . . 8919, 22, 25, 89 , 90, PI. XIII

361759

varianta, Globigerina . . . . . .variata cf. laevis, S piroplectanuninaveslensis, Eponides : . . . . . . .vulgaris. Hoploscaphites constrictus

wilcoxen sis , Alabamina . . . .wilcoxensis, Spiroplectammi na

y

ystadiensis , R osal ina . .ys tadiensi s, Rosalina cf .

11, 2 1, 712072

29

20 , 8416,35

16, 19, 53 , 54 , PI. XV I. . . . . . . . . . S3

PLATES

(Scales correspond to 0.1 mm)

Palaeontologia Polonica, No. 20, 1968

K. PO ZAR YSK A & J. SZCZ ECH U RA: FOR AM INI F ER A F RO M TH E PA L EO C E:" E OF POLAND

PLAT E I

Angulogcrina europaea C USHM AN & EDWARDS

F ig. I. (F. IX/4 ), 215 Ill .

Fig. 2. ( F. IX/5) , 257 111.

Fig. 3. (F . IX /6), 274 ·5111.

Fig. 4 . CF. IX f7), 257111.

Fig. 5. CF . IXIS), 263-8 Ill .

Fig. 6. ( F. IX j9), 274 ·5 Ill.

F ig. 7. (F. IXI] 0), 257 Ill .

All spec imens from Pa m ieto wo boring. 205-274 '5 Ill, Mon tia n

Page

48

Palaeont olouio Polonica. No. ]0 . 1968 PI. I

Palaeontologia Polonica, No. 20, 1968

K. POZARYSKA & J. SZCZ ECH UR A : FORA M IN IFEKA FROM THE PALEOC EN E OF POLAND

PLATE 11

Pa ge

Fig. I. Rosa/ilia brotzeni H OFKER (F. IX/16), 257 Ill . . . 51

Fig. 2. Rosalina koeneni BROTZ EN (F. IX/17), 223 III . . 52

Fig. 3. Discorbis quadrata (TERQUf.M) (F. IXII I) , 257 III . 49

Fig. 4. Rosalina selandiana n. sp., holotype (F. IXfJ8), Nas il6w quarry, 3 m above hard ground layer, Montian , 52

All specimens, except F. IX 11 8, from Pamietowo boring, 205-274'5 m, Montian

Palaeontologia Polonica. No. 10. / 968 PI. Il

~ ....'

4a

K . POZ ARY SK A & .I. SZCZ ECHUKA : FORA MIN IHKA FROM THE PAL EOCENE OF POLAND

Palaeontologia Polonica, No. 20, 1968

K. POZARYSKA & J. SZCZECHU R A : FORAMI NIFERA FROM THE PALEOCEN E OF POLAND

PLAT E 1II

Figs. 1-3. Spirillina er. uodifera TERQUEM (F . IX/22·24), 207 m .

Figs. 4, 5. Patellina sp. (F. IX /20, 21), 207 m . . . . . . . .

Fig. 6. Globulina cf. tubercula ta O'ORIlIG:-iY (F. IX/3), 208·5 m ,

Fig. 7. Globulina sp. (F. IX/2 ), 223 m . . . . . . . . . . . .

Fig. 8. Globulina gibba multlstriata n. sub sp, (F . IX/I ), holotypc, 208·5 m .

Figs. 9-12. Vacuovalvulina ke ijzeri (VAN BELLEN) (F. IX /12-15), 215 m •. .

All specimens from Pamieiowo boring, 205-274,5 m, Montian

58

57

42

42

41

36

Palae ontologia Polonica. No . 10. / 968 PI. 11/

12c

3c3b

12('9b

l Oa ll a 12a

Palaeontologia Polonica, No. 20, 1968

K . POZARYSKA & J. SZCZECHURA : FORAMINIFERA FROM TH E PALEOCENE OF POLAND

PLAT E I V

Figs. 1·3. Rotorbinella mariei ( V AN B ELL EN) (F . IX /47-49), 215 m .

Figs. 4-6. Cuvillierina ? pomeraniana n. sp., holotype (F. IX/25), paratypcs (F. IX/26. 27), 210·7 m

All specimens fro m Pamietowo bori ng, 205-274,5 Ill , Mont ian

Page

54

58

Palaeontologia PI ·o ontca. No. io. 1968 n. IV

Palaeontologia Polonica, No. 20. 1968

K. POZARYSKA & J. SZCZ ECHURA : FORAMI NI FERA FROM THE PALEOCENE OF POLA ND

PLATE V

Figs. I, 3, 4. Rotalia sax orum l)'OKIIIG N Y (F . IX/34), 273·4 m ; (F. IX/35, 36), 208'5 m

Figs. 2, 5 , 7-9. Rotalia marginata n'O RBIGN Y (F. IX/28, 29, 31-33), 208'5 m

Fig. 6. Rotalia alf. marginata O' O RII IG NY (F. IX/30), 208'5 m.

Fig. 10. Rotalia trochidiformis ( L AMARcK) (F. IX/258), 272 m . . . . . . .

All specimens from Pam ietowo bori ng, 205-274'5, Mont ian

Page

61

60

60

62

Palaeontologia Polonica, No. 20, 1968 PI. V

4

8b 8e

5

lOb

lOa

Palaeontologia Polonica, No. 20, 1968

K. POZARYSKA &. J. SZCZ ECH URA : FORA MlN IFE RA FROM TH E PALEOCENE OF POL AND

PLAT E VI

Figs. 1-4. Globorota/ia sp. (F . IX/85, 87, 88), 207 m; (F. IX/86), 231 m .

Figs. 5, 6. Rotorbinella montiana n. sp., holotype (F. IX/50), paratype (F. IX/51), 210·7 m .

Figs. 7-10. Globorota/ia praepseudomenardii HOFKER (F. IX/81-84), 207 m

Fig. 11. Rotorbinella papillata n. sp., holotype (F . IX/52), 210·7 m . . .

All specimens from Pamietowo bori ng, 205-274,5 m, Montian

Page

66

55

66

56

Palaeoutologia PI·• 0 01/1("0 . No . lO. 1968

le 2

PI. VI

Palaeontologia Polonica, No. 20, 1968

K. POZARYSK A & J. SZCZ ECH UR A; FORA MIN IFE RA FROM THE PAL EOC ENE OF POLAND

PLAT E VII

Figs. 1-10. Pararotalia tuberculifera (REUSS) (F. IX/37-41, 43-46), 215 m ; (F. lX/42), 208·5 m

Figs. 11-19. Globorotalia globigeriniformis (V AN BELLEN) (F. IXj65·73), 210·7 m . . . ..

All specimens from Pamietowo boring, 205-274,5 m, Montian

PallC

59

65

Palaeontologia Polonica, No. 20, 1968 PI. VII

[109

11

Palaeontologia Polonica, No. 20, 1968

K. POZAR YSKA & J. SZCZ ECHURA : FORAMIN IFERA FROM T HE PAL EOCENE OF POLA ND

PLATE VIII

Page

Globorotalites granulatus n. sp. . . . . . . . . . . . . . . . 67

Fig. I. Paratype (F. IX/74), 274·5 11\

Figs. 2-5. (F. IX/75-78), 274·5 11\

Fig. 6. Parat ype (F. IX/79), 274·5 m

Fig. 7. Holotype (F. IX/80), Sochaczew boring, 194 11\

All specimens, except F. IX/80, from Pamietowo boring, 205-274,5 m, Montian

Palaeontologia Polonica, No. 20. /9 68 PI. VIIl

Palaeontologia Polonica, No . 20, 1968

K. POZAR YSKA & J . SZCZ ECH URA : FORA MIN IFERA FROM TH E PAL EOCEN E OF POLAND

PLAT E IX

Page

Figs. 1-5. Elphidium cr. lamarcki (O' O RRIGN Y) (F . IX/53-57), 210·7 m . . . . .

Fig. 6. Nonionella communis communis ( D' O RBIGN Y) (F. IX/223), 208'5 m . . .

Figs. 7-9. Nonionella communis paleocenica n. subsp., holotypc (F. IX/226) ; (F.

Figs. 10-12. Nonion granife rum (TERQUEM) (F. IX/220-222), 208·5 m

Figs. 13-15. Nonion sublaeve TES D AM (F. IX/58-60), 208'5 m .

Figs. 16-19. Elphidiella prima (TES D AM) (F . IX/fil-M), 210·7111 .

IX/224, 225) ; 208·5 111

63

82

83

81

81

64

All specimens from Pamictowo boring, 205-274'5 m, Monti an

Palaeontologia Polonica \ '• I O . 20. 1968 PI. IX

I

I

Palaeontologia P%nim. No. 20. 1968

K. POZARYSKA & J. SZCZECH URA : FORAMINI FER A FRO M T HE PALEOCEN E OF POLAND

PLATE X

Figs. I, 2. St omatorbiua sp. (F . IXj254, 255), 231 m . . . . .

Figs. 3-7. Cibicides lectus VASI LENKO (F. IX/105. 109), 231 m .

Figs. 8-12. Cibicides aurouzae R OUVII . I.OIS (F . IX/99-103l. 231 m

All specimens from Pam ietowo bor ing. 205-274'5 Ill. Mont ian

Page

94

77

74

Palaeontologia Polonica, No. 20. 1968 PI. X

lh l e

~113 I I

Palaeontologia Polonica, No . 20, 1968

K. POZAR YSKA & J. SZCZ ECH URA : FORAMI NI FERA FROM THE PALEOCENE OF POLAND

PLAT E XI

Page

Fig . I. Cibicides carinatus (TERQu EM) (F. IX/104), 257 m . . . . . . . . . . . . . . . . . . . . . . 75

Fig. 2. Planulina limbata BROTZEN (F . IX/119), 215 m . . . . . . . . . . . . . . . . . . . . . . . . 80

Figs. 3-7. Cibicides asteroides n. sp. , holotype (F. IX!98), paratypcs (F . IX/96, 97); (F. IX/94-95); 215 m 73

All specimens from Pamictowo boring , 205-274'5 m, Montian

Palaeontologia Polonica, No, ]0. 1968 PI. XI

la

K . POZ ARYSK A & .I. Szn'.~.cHlJ RA: FORAMINIFF RA FROM 'I Ill, P A LEOCEN E OF P OLA ND

Palaeontologia Polonica, No. 20, 1968

K. PO ZAR YSK A & J . SZCZEC HU RA : FORA MIN IF ERA FRO M T HE PALEOC EN E OF POLAND

PLAT E X II

Figs. 1-7. Cibicides cuvillieri R OUVILLOIS (F . IX/ 112-118), 210'7 m .

Fig. 8. Gavelinella lellingensis BR OTZ EN (F. IX/244), 241 m

All specimens from Parnieto wo boring, 205-274'5 m, Montian

Page

76

91

Palaeontolouia Polonica, No . ](). 1968 PI. Xll

K . POZ AR YSK A & J . SJ: CZECHURA: FORAMIN IF ERA IRO\{ T HE P A LEOCE:-lE O F P OLA l"1)

Palaeontologia Polonica. No. 20, /968

K. POZAR YSK A & .I. SZCZ ECH URA : FO RAM IN IFER A '-: RO M TH E PALEOC ENE OF PO LAND

PLAT E XIII

Page

Figs. 1-4. Anomalina ekblomi ( BROTZ EN) (F . IXj245-24ll). 20ll·5 m . . . . . . . . . . . . . . . . . . . . 87

Figs. 5-7. Anomalina umbilicata umbilicata (BROTZEN) (F . IXj238-240). 210·7 m . . . . . . . . . . . . . . . 89

Figs. 8-10. Anomalina umbilicata costata n. subsp., para types (F . IXj24 1. 242). holot ypc ( F. IXj243). 208·5 m 89

All specimens from Parnietowo boring. 205-274'5 m, Montian

Palaeontologia Polonica, No . 20, 1968 PI. X!/l

K. POZ ARYSKA & J. S ZCZECH UR A : FORAM1N IFERA FROM T HE P ALEOCENE Of P OLAND

Palaeontologia Polonica, No. 20, / 968

K. POZAR YSKA & J. SZCZ ECH URA : FOR A MINIFE RA FRO M T HE PALEOC EN E O F POLAND

PLAT E XIV

Figs. 1-5. Anomalina minor n. sp. , paratypes (F. IX/233, 234, 236). holoty pe (F . IXj235), 205 III

Figs. 6-1I. Anomalina danica ( BROTZ EN) (F. IX j227-232), 228 m . . . . . . . . . . . .

All specimens from Pamietowo bori ng, 205-274'5 Ill, Mont ian

Page

87

86

Palaeontologia Polonica, No. 10, /968 PI. X IV

Palaeontologia Polonica, No . 20. 1968

K. POZARYSKA & J. SZCZ ECH URA : FO RA MI [FER A F ROM TH E PALEOCE NE OF POLAND

PLATE XV

Figs. 1-4. Eponides tau/mini (F . IX/89-92), 231 m . . . . . . . . . . . . . . . . . . . . . . . . . .. 72

Fig. 5. Neoeponides cf. schreibersi (D'ORIlIGNY) (F. IX /93), 210'7 m . . . . . . . . . . . . . 72

Figs. 6-10. Mississippina binkhorsti (REUSS) (F. IX/249-251) , 208·5 m, (F. IX /252, 253), 257 m 94

All specimens from Pamietowo boring, 205-274 '5 m, Montian

Palaeontologia Polonica, No. 20, 1968 PI. XV

-

Palaeontologia Polonica, No. 20, 1968

K. POZARYSK A & J. SZCZ ECH URA : FOR AMI NI FER A FR OM T il E PAL EOCEN E OF POL AND

PLAT E XVI

Figs. 1,2. Cibicides proprius (BROTZEN) (F. IX/210, 211), 243 m .

Figs. 3, 4. Cibicides proprius (BROTZEN) (F. IX/147), Bochotnica quarry, Montian

Figs. 5, 6. Anomalina burlingtonensis (JENNINGS) (F. IX/256, 257), 243 m

Fig . 7. Rosalina ystadiensis BROTZ EN (F. IX /19), 257 m . . . . . . . . . . . .

All specimens, except F. IX/147, from Pamietowo boring, 205-274,5 m, Montian

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77

86

53

Pa laeo nto log ia Polonica. No . io . 1961:1 PI. XVI

5b

K. POZARYSKA & J. SZCZ ECH URA : FORAMINIFERA FROM TH E PALEOCENE OF POLAND

Palaeontologla Polonica, No . 20, 1968

K. POZAR YSKA & J. SZCZECH URA : FO RAMINIFE RA FROM TH E PA LEOCENE O F POLAND

PLAT E XVII

Figs . 1-4. Rotorbinella marlei (VAS B ELL EN) (F . IX J259-262)

Figs. 5-8. Rotalia troclridiformis ( L AMARCK) (F. IX/263-266)

All specimens fro m Limburg ( Bunde boring), Montian , ,(GO

54

62

Palaeontologia Polonica, No. 20. 1968 PI. XVII

K. PUZAR YS"; " & J. SZ CU CH UR A : FURAM IN lfmA FRUM THl P ALEOCl Nl or PU LA NlJ

Palaeontologia Polonica, No. 20, 1968

K. POZA RYSK A & J . SZC ZEC HU RA : FO RA MIN IFE RA F RO M T HE PALEOC EN E OF POLA ND

PLAT E XV II I

Figs. 1-4. Rotalia marginata O'ORIlIGN Y (F. IX/267-270)

Figs. 5-9. Rotalia saxorum D ' O RllIGNY (F. IX/271-275)

All specimens from Limburg (Bundc boring), Montian . .( (OC'

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61

Pa laeontologia Polonica . No . io. 19f>S PI. XVIII

K . POZARYS" A & J. SZCI.EC Il URA : FORAMIN IHiRA I'RUM rilE P ALH JCENE 0 1 POLAND